Description from Flora of China
Thelypteris sect. Parathelypteris H. Itô in Nakai & Honda, Nov. Fl. Jap. 4: 127. 1939; Coryphopteris Holttum; Wagneriopteris A. Löve & D. Löve.
Plants small to medium-sized, terrestrial plants, rarely in marshes or meadows. Rhizomes long creeping or short and decumbent, ascending or erect, glabrous or with scales or rusty yellow hairs. Fronds remote, approximate, or clustered; stipes stramineous or castaneous, ± polished, bases sometimes nearly black and glabrous or with spreading, grayish white, multicellular acicular hairs, distally glabrous or pubescent; laminae ovate-oblong, oblong-lanceolate, or lanceolate, tapering or not tapering to bases, pinnate-pinnatifid, acuminate and pinnatifid at apices; lateral pinnae mostly narrowly lanceolate to linear-lanceolate, bases symmetrical, truncate or broadly cuneate, not adnate to rachises, sessile or occasionally shortly stalked, apices acuminate; proximal pinnae not shortened or 1 to several pairs of pinnae obviously shortened, even reduced to small auricles, pinnatifid; segments mostly oblong, rectangular, or subsquare, margins entire or ± serrate, apices rounded-obtuse, sometimes pointed or with sinuslike angles. Veinlets pinnate, free, simple and all reaching margins. Laminae herbaceous or papery, when dry yellowish green, sometimes brownish green or nearly black, both surfaces ± with pubescent or acicular hairs, rarely glabrous when dry, sometimes with orange-yellow or reddish purple glands abaxially; costae grooved adaxially, densely covered with short setae, rounded and raised abaxially and usually ± with acicular hairs, rarely glabrous. Sori orbicular, medium-sized, dorsifixed at middle or near ends of veinlets, located between main veins and margins or slightly closer to margins; indusia larger, orbicular-reniform, sometimes horseshoe-shaped, when dry brown, membranous, glabrous or hairy, usually persistent. Spores bilateral, orbicular-reniform, perispores thin and transparent, corrugate, ± finely reticulate on surfaces. x = 8, 9, 31.
Parathelypteris is very similar to Metathelypteris in shape, but it differs by the costae grooved adaxially; the veinlets usually simple, not forked, and reaching margins; and the indusia larger and readily discerned.
The following taxa are excluded from the present treatment, pending further research: Parathelypteris auriculata H. G. Zhou & H. Li (Acta Bot. Yunnan. 14: 33. 1992), described from Guangxi (Wuming), and P. jinfoshanensis Ching & Z. Y Liu (Bull. Bot. Res., Harbin 4(3): 18. 1984), described from Sichuan (Nanchuan).
About 60 species: tropical and subtropical regions of E Asia, SE Asia to Pacific islands; 24 species (11 endemic) in China.
(Authors: Lin Youxing (林尤兴); Kunio Iwatsuki)