Herbs perennial, tuberous. Leaves mostly near middle of stem and distal; leaf blade deeply palmately divided; primary lobes 2- to many lobulate. Petals of same color as sepals, not black-brown, distally not enlarged. Staminodes of same color as sepals, not black-brown, distinctly differentiated into a claw and an expanded limb; limb abaxially yellow barbate. Carpels 3. Seeds transversely squamulose winged.

About 45 species: N Africa, W Asia, S Europe, North America; one species in China.