3. Leucophanes Brid., Bryol. Univ. 1: 763. 1827.
Plants soft, slender, grayish green to yellow or glaucous green, in loose to dense tufts. Stems erect, orange-brown to dark red, sparsely branched; rhizoids papillose. Leaves erect to more or less contorted when dry, loosely overlapping when moist, narrowly to broadly lanceolate, concave to keeled at base or through most of their length, plane above, acuminate, acute, obtuse, retuse or bent backwards at the apex; margins serrate particularly in the upper third with single or paired teeth; marginal stereome forming a distinct, sharply differentiated border; basal hyaline lamina asymmetrical, extending to 1/5 – 1/3 leaf length on both sides of costa, irregularly bistratose, cells linear; chlorocysts in cross section quadrangular; costa percurrent to shortly excurrent, smooth to strongly papillose on back and tips, convex on the abaxial side, flat on the adaxial side, in cross section, central stereid band present. Gemmae on leaf tips, papillose, oblong to clavate, 6–10 cells long. Dioicous or monoicous. Sporophytes rare; setae 3.1–15 mm long, sinistrorse, smooth; capsules 1.8–3.0 mm long, dark orange, erect to horizontal; stomata superficial at neck of urn; peristome teeth 16, more or less papillose, lanceolate, with a colorless properistome, 2–3 cells high; opercula long, subulate-rostrate. Calyptrae cucullate. Spores 10–20 µm in diameter.
The genus Leucophanes consists of 25 species in the world, mainly with tropical distributions. Three species are reported from China, but only two species are recognized in this study. Leucophanes candidum (Schwaegr.) Lindb. was reported from Hainan, Taiwan, and Yunnan. All these records turned out to be Leucophanes octoblepharioides Brid. Leucophanes is a leucobryoid moss genus with chlorocyst and leucocyst structures similar to those of Leucobryum. However, the peristome structure of Leucophanes is of the type found in the Calymperaceae.