10. Leucobryum scabrum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi. 2: 292. 1966.
Plants medium-sized to large, up to 5 cm high, whitish to grayish green, in dense, compact tufts. Stems erect, simple or branched; central strand absent. Leaves erect-spreading, 5–8 mm long, lanceolate, gradually narrowed to channelled apices from broadly ovate to oblong-ovate base, acute at the apex; dorsal side of leaf acumina undulate and spinosely prorate; laminae bordered by 1 (–2) rows of narrowly rectangular cells, laminae near leaf base consisting of 5–6 rows of narrowly rectangular cells, basal marginal cells indistinct; costae in cross section consisting of a median layer of quadrangular chlorocysts with 1–3 layers of leucocysts on either sides. Dioicous. Sporophytes not seen. [According to Yamaguchi (1993), setae 2.5–3.2 cm long; capsules inclined, ovoid to ellipsoid, 1.6–1.8 mm long. Other sporophytic features were reported to be the same as those described under the generic description.]
Type. Japan: Siebold s.n.
Chinese specimens examined: ANHUI: Huang Shan (Mt.), P.-C. Chen et al. 7288 (IBSC, PE). FUJIAN: De-hua Co., P.-Z. Zheng 567 (IBSC); San-ming City, D.-K. Li 01490 (SHM). GUANGDONG: Bo-luo Co., N.-H. Mou 1, 3 (IBSC, SYS); Cong-hua Co., H. Wu 507 (IBSC); Ding-hu Shan (Mt.), H. Wu 191 (IBSC); Lian-ping Co., P.-J. Lin 1336(a) (IBSC); Ru-yuan Co., P.-J. Lin 615 (IBSC); Xin-xing Co., P.-J. Lin 490 (IBSC); Ying-de Co., X.-H. Xu 11(2) (IBSC). GUANGXI: Shang-si Co., P.-J. Lin & C. Gao 1969 (IBSC, IFSBH). HAINAN: Jie-feng-ling (Mt.), P.-C. Chen et al. 375 (IBSC, PE). JIANGXI: Jing-gang Shan (Mt.), D.-K. Li 16743 (IBSC, SHM); Lu Shan (Mt.), D.-K. Li 017025 (IBSC, SHM). SICHUAN: Chuangqing City, W.-M. Zhu 7701 (IBSC, YUKU); Omei Shan (Mt.), W.-M. Zhu 061 (IBSC, YUKU). TAIWAN: Tao-yuen Co., M.-J. Lai 0233 (MO, NICH). YUNNAN: Ping-bian Co., W.-M. Zhu 708b (IBSC, YUKU). ZHEJIANG: Long-quan Co., P.-C. Wu 471 (IBSC, PE).
Habitat: on soil and tree trunks under forests; Distribution: China, Japan, Malaysia, and Thailand.
Illustrations: Pl. 68, figs. 4–6. Yamaguchi 1993 (Pls. XI–XIV).