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Moss China | Family List | Moss China V. 2 | Fissidentaceae | Fissidens

21. Fissidens grandifrons Brid., Muscol. Recent. Suppl. 1:170. 1806.


Fissidens diversiretis Broth., Symb. Sin. 4: 11. 1929. Type. China: Yunnan, between Ngaiwa and Mekong, 7 X 1915, Handel-Mazzetti 8477 (holotype H-BR). Fissidens grandifrons var. planicaulis (Besch.) Nog., J. Hattori Bot. Lab. 7: 68. 1952. Fissidens planicaulis Besch., Ann. Sci. Nat. Bot., sér. 7, 17: 335. 1893. Fissidens yunnanensis Besch., Rev. Bryol. 18: 88. 1891. Type. China: Yunnan, Delavay 4631 (isotype H-BR).

Plants medium-sized to large, creeping, dark green, rigid, brownish when old. Leafy stems simple or branched, 16–86 mm long, 2.5–3.0 mm wide; axillary hyaline nodules differentiated; cortical cells small, thick-walled; central strand not differentiated. Leaves in 13–83 pairs, densely arranged, stiff even when dry, the lowest leaves small, middle to upper leaves lanceolate to ensiform-lanceolate, 2.8–3.5 mm × 0.4–0.5 mm, obtuse to acute at apex; base of dorsal laminae wedge-shaped, decurrent; vaginant laminae about 1/2 the leaf length, equal; costa stout, but obscure, ending a few cells below leaf apex; margins slightly serrulate; in cross section apical and dorsal laminae 1–2 cells thick at margin, 3–6 cells thick near costa, one cell thick for most parts of vaginant laminae; laminal cells near margin smaller, with thinner cell walls, larger, with thicker cell walls toward costa; cells of apical and dorsal laminae quadrate to hexagonal, 7–11 µm long, smooth, moderately thick-walled to thick-walled; cells of vaginant laminae similar to those of apical and dorsal laminae, but larger, with thicker walls toward base. Dioicous. Female inflorescences axillary. Archegonia 400–640 µm long, with stalks up to 140 µm long. Setae 18–21 µm long, smooth; capsules erect to horizontal, symmetrical; urns cylindric, 1.1–1.6 mm long; exothecial cells quadrate to shortly rectangular; peristome teeth ca. 0.43 mm long, 114 µm wide at base. Spores 14–25 µm in diameter.

Type. U.S.A.: “Nova Anglia.”.

Chinese specimens examined: GUANGXI: Long-sheng Co., P.-C. Wu & Y.-X. Lin 1211 (PE, SYS). GUIZHOU: Sui-yang Co., B.-C. Tan 91-989 (FH, MO). HUBEI: Sheng-nong-jia, P.-C. Wu 231 (MO, PE), Sino-American Team 1670 (H, MO, NY, PE). QINGHAI: Nang-qen Co., B.-C. Tan 95-1681 (FH, MO); Yu-shu Co., B.-C. Tan 95-1711 (MO). SICHUAN: Du-kou City, M.-Z. Wang 20118b (PE, SYS); Tian-quan Co., S. He 32050 (MO). TAIWAN: Hua-lien Co., Shevock 14435 (MO); Nan-tou Co., M.-J. Lai 2788 (H); Ping-tung Co., M.-J. Lai 9850 (H, MO), C.-C. Chuang 1571 (MO); Tai-zhong Co., M.-J. Lai 8699 (H, MO). XIZANG: Jia-yu-zhunba, M. Zang 987 (HKAS, SYS); Ri-tu Co., D.-D. Tao 5308c (IBSC, SYS). YUNNAN: Li-jiang Co., M. Zang 1289 (HKAS, SYS); Yang-bi Co., Redfearn et al. 234 (MO).

Habitat: on wet or submerged rocks; Distribution: China, Korea, Japan, Nepal, India, North and Central Africa, and North and Central America.

Illustrations: Z.-H. Li 1985 (Fig. 28, a–m).


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