50. Fissidens tosaensis Broth., Öfvers. Förh. Finska Vetensk.-Soc. 62A(9):5. 1921.
Fissidens hetero-limbatus Sak., Bot. Mag. (Tokyo) 47: 736. 1933.
Plants small, green to brownish. Leafy stems usually simple, 2.6–5.0 mm long, 1.8–3.5 mm wide; axillary hyaline nodules indistinctly differentiated; central strand weakly differentiated. Leaves in 4–9 pairs, middle to upper leaves oblong-lanceolate to ovate-lanceolate, 1.3–1.6 mm × 0.4–0.5 mm, acute at apex; base of dorsal laminae rounded to wedge-shaped, rarely slightly decurrent; costa strong, percurrent to shortly excurrent; margins nearly entire except at the leaf apex, which is often slightly serrulate; vaginant laminae ca. 1/2 –3/5 the leaf length, equal or slightly unequal; limbidia strong, 1–2 rows of cells on apical laminae, 2–5 rows of cells on vaginant laminae, 1–3 cells thick in cross section; cells of apical and dorsal laminae quadrate to hexagonal, 7–14 µm long, moderately thick-walled, smooth; cells of vaginant laminae similar to those of apical and dorsal laminae, but larger and longer toward base near costa. Autoicous. Male inflorescences bud-like, often growing in axils of stem leaves. Female inflorescences mostly in axils of stem leaves, occasionally terminal on stems. Perichaetial leaves not markedly differentiated. Archegonia 160–210 µm long. Setae 6.0–8.3 mm long, smooth; capsules curved, asymmetrical; urns 0.6–0.8 mm long; opercula long rostrate, ca. 0.5 mm long; exothecial cells quadrate to rectangular, thin-walled to partially thick-walled, with thick-walled corners; peristome teeth linear, 0.2–0.3 mm long, 38 µm wide at base. Calyptrae mitrate, ca. 0.5 mm long. Spores 7–12 µm in diameter.
Type. Japan: Prov. Tosa, S. Okamura 29 (holotype H-BR).
Chinese specimens examined: FUJIAN: Wuyi Shan (Mt.), P.-J. Lin 2735 (IBSC, SYS). GANSU: Wen Co., Z.-P. Wei 6847 (HKAS, NICH). GUANGDONG: Shi-xing Co., P.-J. Lin & Y.-Y. Yang 4620 (IBSC, SYS). HAINAN: Ba-wang-ling (Mt.), 1974 Team 3281 (IBSC, SYS). JIANGSU: Nan-jing City, M. Zang 1001 (HKAS, NICH). SHAANXI: Hu Co., M. Wang 469 (HKAS, NICH). SICHUAN: Cheng-du City, J.-L. Yang M008 (IBSC, SYS). YUNNAN: An-ning Co., Koponen 37905 (H. SYS); Gong-shan Co., M.-Z. Wang 9704 (PE, SYS); Gao-li-gong Shan (Mt.), M.-Z. Wang 9345c (PE, SYS); Jing-hong Co., Redfearn et al. 34042 (MO); Kun-ming City, Koponen 37424 (H, SYS); Wei-deng, M.-Z. Wang 6496d (PE, SYS); Yang-bi Co., Redfearn et al. 163 (MO). ZHEJIANG: Pu-tuo Shan (Mt.), D.-K. Li 6935 (SHM, SYS).
Habitat: on soil; Distribution: China and Japan.
Illustrations: Pl. 89, figs. 1–17.