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Pakistan | Family List | Labiatae

12. Dracocephalum L., Sp. Pl. 594. 1753. Gen. P1. ed. 5: 258. 1754; Benth. in DC., Prodr. 12: 396. 1848; Hook. f., Fl. Brit. Ind. 4: 664. 1885; Briquet in Engler & Prantl, Nat. Pflanzenfam. ed. 1, 4, 3A: 238. 1896; Mukerjee in Rec. Bot. Surv. Ind. 14, 1: 135. 1940; Schischkin in Komarov, Fl. URSS 20: 439. 1954; Rech. f., Fl. Iran. 150: 218. 1982; Budantzev in Bot. Zhum. 72, 2: 260-267. 1987.

I.C. Hedge

  • Moldavica Adans.

    Herbaceous or suffruticose perennials, rarely annuals. Stems leafy, much branched, pubescent with simple hairs or almost glabrous. Leaves not divided to pinnatisect, petiolate. Flowers in verticillasters borne in axils of upper leaves or in terminal oblong or capitate heads. Bracts entire or lobed, aristate or not. Flowers white, pink, blue to violet. Calyx tubular-campanulate, 15-nerved, straight or curved, bilabiate or scarcely so; upper lip equally 3-toothed or middle lobe dearly larger than laterals; lower lip with 2 teeth or apparently 4; base of teeth always with swollen folds at sinuses; tube glabrous within. Corolla bilabiate; tube gradually widening from base; upper lip straight or curved, subentire or bifid without longitudinal folds; lower lip 3-lobed with median clearly larger. Stamens 4, usually included under corolla upper lip, rarely exserted, didynamous with the posterior (upper) stamens slightly longer than the anterior (lower); thecae pilose or glabrous, bilocular, divergent. Style with 2 almost equal lobes. Nutlets 4, ovoid-oblong, smooth, weakly trigonous, apically truncate-rounded, frequently mucilaginous on wetting.

    About 40 mainly Eurasian species; some are high alpines. Very closely related to Nepeta, but calyces with a thickened fold at the sinuses between the teeth; also close to Lallemantia.

    Species doubtfully in our area

    Dracocephalum aitchisonii Rech. f. (Kongel Danske Vidensk. - Selsk. Skr. 8, 1: 42. 1954) is recorded by Rechinger in Flora Iranica (150: 228. t. 213. 1982) from Pakistan, Kurram valley, based on Aitchison 573, but the localities on the field notes given by Aitchison are all on the Afghan side of the frontier, at highish altitudes around Mt Sikaram. However, the species may yet be found in Pakistan; it is recognized by the perennial habit, the deeply lobed leaves, the ± capitate inflorescences, the prominent median lobe of the upper calyx lip, the conspicuous thickening at the calyx sinuses and the violet corollas.


    1 Upper lip of calyx with a broad ovate median lobe clearly larger than laterals; calyx distinctly bilabiate   (2)
    + Upper lip of calyx subequally 3-lobed; calyx bilabiate or scarcely so   (3)
           
    2 (1) Caespitose perennial with stems upto 5 cm; leaf margins deeply crenate to pinnatifid   2 Dracocephalum paulsenii
    + Erect perennial with stems upto 30 cm; leaves ± regularly crenulate   3 Dracocephalum nutans
           
    3 (1) Stamens long-exserted; leaves broad ovate to orbicular; lowermost leaves with petioles upto 33 cm   1 Dracocephalum stamineum
    + Stamens included; leaves ovate to oblong; petioles less than 2 cm   (4)
           
    4 (3) Annual; calyx 8-10 mm   6 Dracocephalum moldavica
    + Perennial; calyx 12-18 mm   (5)
           
    5 (4) Leaves ± regularly crenate, Cordate; corolla white tinged pink or lilac   5 Dracocephalum heterophyllum
    + Leaves pinnatisect to 1-pinnate; Corolla violet-blue or mauve   4 Dracocephalum bipinnatum

    Lower Taxa

    Related Synonym(s):


     

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