Barbula sect. Aloina J. K. A. Müller
Plants in thin turfs or scattered. Stem 0.5--6 mm, central strand present. Leaves forming a rosette, lingulate, suborbicular or deltoid, deeply concave, 0.5--2.8 mm, infolded, partly 2-stratose, base auriculate, sheathing, margins entire to irregularly crenulate distally, apex cucullate, mucronate or piliferous; costa broad, percurrent to long-excurrent, in section hydroids absent, stereid band abaxial, occasionally wanting, filaments present on costa and adjacent 2-stratose area of leaf blade, filament cells cylindrical to subspheric, thin-walled, terminal cell subspheric to nearly conic, with an apical thickening; cells of leaf base thin-walled, rectangular, sometimes differentiated marginally, medial and distal leaf cells thick-walled, mostly transversely elongated, smooth. Sexual condition monoicous or dioicous, perigonia distinct, with non-cucullate unfolded leaves, perichaetia sometimes partly differentiated, the inner leaves with unfolded blades. Seta 4.5--17 mm, capsule cylindric to ovoid-cylindric, erect or slightly inclined, annulus of 1--2 rows of vesiculose cells, deciduous or persistent, operculum conic to long-rostrate, erect or inclined, peristome generally long and twisted, papillose, with a basal membrane. Calyptra cucullate. Spores spheric, finely papillose, 9--25 µm.
Species 9 (5 in the flora): mostly on calcareous soils in the Mediterranean Basin, southern Africa, Australia; sparsely distributed from Alaska to Argentina.
This is generally a very distinct genus characterized by differentiated leaf bases, infolded partly 2-stratose laminae, broad costae, and filaments ending in a terminal cell with an apical thickening; the filaments cover the costa and part of the lamina. Plants of Aloina sometimes resemble Indusiella (Grimmiaceae) in habit and general leaf structure although no filaments are found on the leaves of the latter.
SELECTED REFERENCES
Delgadillo M., C. 1973. A new species, nomenclatural changes, and generic limits in Aloina, Aloinella, and Crossidium. Bryologist 76: 271--277. Delgadillo M., C. 1975. Taxonomic revision of Aloina, Aloinella and Crossidium (Musci). Bryologist 78: 245--303.
