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Takakia ceratophylla (Mitten) Grolle, Oesterr. Bot. Zeitschr. 110(4): 444, f. 1. 1963.
Authors: John R. Spence & W. B. Schofield
Lepidozia ceratophylla Mitten
Plants lacking distinctive odor when dry, distal portion of leafy shoot readily caducous. Stems in cross section with 2 layers of thicker walled epidermal cells, stoloniferous stems mainly subterranean. Leaves typically of thick-walled cells, in 4 segments, connate at base, cross section of each segment with 3--5 inner cells and 10--15 smaller outer epidermal cells, these outer cells small, 25 × 10 µm, regularly arranged. Slime hairs thick-walled. Capsule terminal, usually single.
Capsules mature late summer--early fall (July--Sept.). Uncommon to rare on moist soil and shaded banks, slopes and over rocks in tundra; 75--700 m; Alaska; Asia (China in Yunnan, India in Sikkim, Nepal).
This species appears to be somewhat tolerant of desiccation. Frequently it grows with other drought-tolerant bryophytes such as Andreaea and Gymnomitrion. The shoots are more rigid than those of T. lepidozioides and the leaf arrangement more regular. Rhizomatous shoots are predominantly subterranean.
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