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8. Gleicheniaceae C. Presl

Forking Fern Family

Clifton E. Nauman

Plants coarse, terrestrial. Stems long-creeping, forked, stele protostelic (a solid rod of vascular tissue with phloem surrounding xylem), covered with scales or hairs. Leaves monomorphic, large, scrambling or trailing, 1--many times forked. Petiole not articulate to stem, with pair of opposite pinnae and arrested bud at apex, or rachis continuing and producing 2 or more pairs of opposite pinnae. Pinnae 1--several times forked, with arrested bud at each fork; indument of simple, branched, or stellate hairs [scales]. Veins free, 1--4-forked. Sori round, indusia absent. Sporangia 2--many on slightly elevated receptacle, sessile to subsessile; annulus complete, transverse, medial, longitudinally dehiscent; spores 120--800 per sporangium. Spores all alike, whitish to yellowish, bilateral to globose, monolete or trilete, generally smooth without elaborately ornamented surface. Gametophyte borne aboveground, green, obcordate to elongate.

Genera 4, species ca. 140 (1 genus, 1 species in the flora): nearly worldwide in tropical to subtropical regions.


Holttum, R. E. 1957. Morphology, growth habit, and classification in the family Gleicheniaceae. Phytomorphology 7: 168--184. Maxon, W. R. 1909. Gleicheniaceae. In: N. L. Britton et al., eds. 1905--1972. North American Flora. New York. Vol. 16, pp. 53--63. Underwood, L. M. 1907. American ferns --- VIII. A preliminary review of the North American Gleicheniaceae. Bull. Torrey Bot. Club 34: 243--262.

Lower Taxon


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