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FNA | Family List | FNA Vol. 8 | Ericaceae

38. Lyonia Nuttall, Gen. N. Amer. Pl. 1: 266. 1818.

Maleberry, staggerbush [For John Lyon, 1765-1814, Scottish-born, early American botanist and explorer of southern Appalachians]

Walter S. Judd

Arsenococcus Small; Desmothamnus Small; Neopieris Britton

Shrubs or trees, (sometimes with woody burl, resprouting after fire). Stems erect to arching; twigs hairy, sometimes glabrescent (sometimes also lepidote). Leaves deciduous or persistent; blade elliptic, obovate, or ovate, membranous to coriaceous, margins entire, undulate, or serrulate [irregularly serrate], plane or revolute, surfaces multicellular, peltate-scaled or short-stalked stipitate-glandular-hairy, often unicellular-hairy on major veins or abaxial surface; venation brochidodromous (or reticulodromous). Inflorescences axillary fascicles, panicles, or racemes, (2-)5-8(-12)-flowered, sometimes flowers solitary, (produced just before flowering). Pedicels: bracteoles 2, at or near base. Flowers: sepals usually [4-]5[-8], slightly connate, ovate-deltate to lanceolate; petals usually [4-]5[-8], connate nearly their entire lengths, white to red, corolla cylindric to urceolate, lobes much shorter than tube, (sparsely stipitate-glandular-hairy or peltate-scaled); stamens usually [8-]10[-16], included; filaments geniculate, flattened, roughened or hairy, with or without spurs; anthers without awns, dehiscent by elliptic pores; pistil usually [4-]5[-8]-carpellate; ovary [4-]5[-8]-locular; (style slightly longer than stamens); stigma capitate-truncate. Fruits capsular, globose to ovoid or ellipsoid, dry, (with [4-]5[-8] pale, decidedly thickened, whitish sutures). Seeds ca. 100-300, ellipsoidal, narrowly oblong, obovoid to angular-obovoid or narrowly conic, (sometimes tailed); testa cells elongate. x = 12.

Species 36 (5 in the flora): e United States, Mexico, West Indies, e Asia.

SELECTED REFERENCES Judd, W. S. 1981. A monograph of Lyonia (Ericaceae). J. Arnold Arbor. 62: 63-209, 315-436. Judd, W. S. 1995. Lyonia. In: Organization for Flora Neotropica. 1968+. Flora Neotropica. 98+ nos. New York. No. 66, pp. 222-294.

1 Abaxial surface of leaf blades, pedicels, and calyx lepidote   (2)
+ Abaxial surface of leaf blades, pedicels, and calyx nearly glabrous or hairy, not lepidote   (3)
2 (1) Ultimate branchlets spreading, flowers nearly always restricted to previous year’s branches; leaves not conspicuously reduced toward branch tips, blade margins usually revolute distally, sometimes strongly so, major veins usually depressed; shrubs or trees to 6(-12) m.   4 Lyonia ferruginea
+ Ultimate branchlets rigidly ascending, flowers frequent on current year’s branchlets (some flowers also borne on branches of previous year); leaves conspicuously reduced toward branch tips, blade margins at most slightly revolute distally, major veins not depressed; shrubs to 1.5(-3) m.   5 Lyonia fruticosa
3 (1) Leaf blades scattered, multicellular long-headed-hairy abaxially, margins serrulate; corollas urceolate, 2-4(-4.5) mm; calyx lobes 0.5-1.5 mm.   1 Lyonia ligustrina
+ Leaf blades scattered, multicellular short-headed-hairy abaxially, margins entire; corollas cylindric, 5-14 mm; calyx lobes 2-9.5 mm   (4)
4 (3) Leaves persistent, blade rigidly coriaceous, intramarginal vein present; corollas usually pink, sometimes red or white, 2-5 mm wide, base swollen; filaments roughened; capsules ovoid to ovoid-globose   2 Lyonia lucida
+ Leaves deciduous, blade membranous, intramarginal vein absent; corollas usually white, rarely pink, 4.5-9 mm wide, base not swollen; filaments long-hairy, especially near base; capsules ovoid.   3 Lyonia mariana

Lower Taxa


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