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FOC | Family List | FOC Vol. 14 | Ericaceae

18. Lyonia Nuttall, Gen. N. Amer. Pl. 1: 266. 1818, nom. cons., not Rafinesque (1808).

珍珠花属 zhen zhu hua shu

Fang Ruizheng (方瑞征 Fang Rhui-cheng); Peter F. Stevens

Xolisma Rafinesque.

Shrubs or trees, deciduous or evergreen, often from underground woody burl or producing thickened horizontal underground rhizomes. Buds flattened, conical or ovoid, usually with 2 large imbricate glabrous scales. Leaves spirally arranged, petiolate; leaf blade entire [toothed in United States]. Inflorescences axillary, racemose. Flowers [4- or]5[–8]-merous. Calyx with variously estivated lobes, usually valvate in bud. Corolla white [to red], tubular or urceolate; lobes short. Filaments flattened, geniculate, with or without 1 pair of spurs at anther-filament junction; anthers dehiscing by introrse-terminal elliptic pores. Ovary superior, with many ovules per locule; stigma truncate. Capsule loculicidal, with pale ± thickened sutures sometimes separating from valves in dehiscence. Seeds oblong-ovoid or spindle-shaped to shortly linear, minute, ends often truncated.

Thirty-five species: E Asia, North America; five species (two endemic) in China.

1 Capsule with slightly thickened sutures; leaves 1.5–7 × 0.6–4 cm, apex obtuse, mucronate.   (2)
+ Capsule with much thickened sutures; leaves 4–20 × 2.5–12 cm, apex acuminate to abruptly acuminate.   (3)
2 (1) Filaments with 2 spurs near apex; inflorescences 6–15 cm; rachis sparsely to densely pubescent; ovary pubescent.   4 Lyonia compta
+ Filaments without spurs; inflorescences 1–4(–7) cm; rachis densely fulvous-pubescent; ovary glabrous to pubescent.   5 Lyonia villosa
3 (1) Calyx lobes 6–11 mm; filaments without spurs; inflorescences 2–7 cm; leaf blade usually leathery.   3 Lyonia macrocalyx
+ Calyx lobes 2–6 mm; filaments with 2 spurs near apex; inflorescences 5–20 cm; leaf blade usually papery.   (4)
4 (3) Rachis of inflorescences, pedicels, and abaxial surface of calyx pubescent; leaf blade apically usually acuminate.   1 Lyonia ovalifolia
+ Rachis of inflorescences, pedicels, and abaxial surface of calyx glabrous; leaf blade apically usually abruptly acuminate.   2 Lyonia doyonensis

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