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45. Sida Linnaeus, Sp. Pl. 2: 683. 1753; Gen. Pl. ed. 5, 306. 1754.
Fanpetals, wireweed [Greek side, name used by Theophrastos for plants now called Nymphaea alba Linnaeus] Fanpetals, wireweed [Greek side, name used by Theophrastos for plants now called Nymphaea alba Linnaeus]
Paul A. Fryxell Steven R. Hill
Dictyocarpus Wight; Malvinda Boehmer; Pseudomalachra (K. Schumann) Monteiro
Herbs, annual or perennial, subshrubs, or shrubs. Stems erect, ascending, or reclining to procumbent, glabrous or hairy, sometimes viscid (S. glabra). Leaves spirally arranged (distichous in S. planicaulis and S. ulmifolia), petiolate or subsessile; stipules persistent, usually linear to lanceolate or falcate; blade usually unlobed (lobed with maplelike leaves in S. hermaphrodita), base cuneate, cordate, subcordate, truncate, or rounded, margins crenate, dentate, serrate, or entire. Inflorescences axillary solitary (sometimes paired or clustered) often plicate in bud, usually 1/2 divided, often 10-ribbed at base (unribbed in S. hermaphrodita) or angulate, lobes acute or acuminate to triangular or ovate; corolla white, cream, yellow, yellow-orange, salmon-pink, red-orange, or reddish [purplish], sometimes with dark-red center; staminal column included; style 5–14-branched; stigmas capitate. Fruits developed or muticous, reticulate, glabrous or hairy, lateral walls usually persistent, indehiscent below with well-differentiated dorsal wall, indehiscent or partially dehiscent apically. Seeds 1 per mericarp, glabrous. x = 7, 8.
In the flora area, Sida linifolia Cavanilles, flax-leaved sida, is known from a single collection (Alabama, Mobile, introduced from West Indies on ballast, Sep 1886, Mohr s.n., F) and treated here as a waif; it is distinguished from other sidas in North America by its entire leaf margins. Sida cordata (Burman f.) Borssum Waalkes has been reported in Maryland (Baltimore City); it is a generally prostrate herb with cordate leaves and filiform pedicels that are nearly the same length as the leaves; no vouchers have been found; if it was present, it can be regarded as a waif. Reports of S. aggregata K. Presl, a variable and rather common Neotropical species, have not been verified; no vouchers have been located. Sida acuta Burman f. and S. carpinifolia Linnaeus f. are names often used for ballast specimens of plants found in temperate seaports that have not persisted. Most sidas have apical spines on the fruits that adhere to fur, wool, and clothing, and therefore it may be difficult to pinpoint their native ranges versus the areas to which they have been introduced. Some are considered to be pan-tropical roadside weeds.
SELECTED REFERENCES Fryxell, P. A. 1985. Sidus sidarum V. The North and Central American species of Sida. Sida 11: 62–91. Kearney, T. H. 1954. A tentative key to the North American species of Sida L. Leafl. W. Bot. 7: 138–150. Krapovickas, A. 2003b. Sida sección Distichifolia (Monteiro) Krapov. comb. nov., stat. nov. (Malvaceae–Malveae). Bonplandia (Corrientes) 12: 83–121. Krapovickas, A. 2007. Las especies de Sida secc. Malachroideae del Cono Sur de Sudamérica. Bonplandia (Corrientes) 16: 209–225. Siedo, S. J. 1999. A taxonomic treatment of Sida sect. Ellipticifolia (Malvaceae). Lundellia 2: 100–127.
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| 1 |
Leaf blades palmately 5–7-lobed, maplelike, to 24 cm; calyces not ribbed or angled. |
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7 Sida hermaphrodita |
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Leaf blades unlobed, 1–9 cm; calyces ribbed or angled |
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(2) |
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| 2 (1) |
Stems procumbent |
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(3) |
| + |
Stems erect, sometimes ascending to reclining but not procumbent |
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(4) |
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| 3 (2) |
Petals white; leaves distributed evenly along stems, blade margins crenate to base; stems usually with 1–2 mm simple hairs in addition to multirayed stellate hairs; mericarps slightly rugose. |
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1 Sida abutilifolia |
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Petals salmon-pink, red-orange, or yellowish; leaves crowded distally on stems, blade margins dentate only at apex; stems with appressed, normally 4-rayed stellate hairs; mericarps prominently muricate. |
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3 Sida ciliaris |
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| 4 (2) |
Styles 5-branched; mericarps 5; leaf blades cordate or subcordate at base |
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(5) |
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Styles 7–14-branched; mericarps 7–14; leaf blades usually cuneate or truncate at base, sometimes rounded or subcordate to cordate |
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(7) |
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| 5 (4) |
Stems and petioles minutely stellate-hairy, hairs to 0.5 mm; petioles usually with small spinelike tubercle on stem just below its attachment. |
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16. Sida spinosa |
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Stems and petioles glandular-viscid and/or with simple hairs 1–3 mm; petioles without spinelike tubercle just below attachment to stem |
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(6) |
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| 6 (5) |
Stems usually glandular-viscid; petals white or yellow-orange without darker base, not fading rose-pink; calyces usually glandular, not setose, lobes triangular, acute to short-acuminate (not beaked in bud). |
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6 Sida glabra |
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Stems not glandular-viscid; petals yellowish to orange or salmon usually with dark orange or reddish base, fading rose-pink; calyces setose, not glandular, lobes trullate, attenuate-aristate (beaked in bud). |
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19 Sida urens |
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| 7 (4) |
Leaves and branches distichous |
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(8) |
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Leaves spirally arranged |
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(9) |
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| 8 (7) |
Mericarps 7 or 8, spines 2 mm; inflorescences usually axillary glomerules, sometimes flowers solitary or paired; calyces 5–6 mm; staminal columns glabrous. |
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12 Sida planicaulis |
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Mericarps 8–12, spines 0.5–1 mm; inflorescences axillary, flowers solitary or paired; calyces 6–8 mm; staminal columns hairy, sometimes glabrous. |
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18 Sida ulmifolia |
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| 9 (7) |
Leaf blades ± ovate or ovate-oblong with broadly cordate base, infrequently ovate-lanceolate to lanceolate; petioles (5–)10–25 mm |
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(10) |
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Leaf blades rhombic to subrhombic or elliptic to oblong, sometimes oblong-lanceolate, lanceolate-elliptic, round, lanceolate, or linear; petioles 2–10(–40) mm |
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(11) |
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| 10 (9) |
Leaf blades velvety-tomentose, 1–2 times longer than wide; calyces prominently ribbed, densely stellate-tomentose; inflorescences axillary, usually corymbs or panicles, sometimes solitary flowers; fruits 6–7 mm diam.; mericarp spines to 2 mm; Alabama, Florida, Texas. |
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4 Sida cordifolia |
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Leaf blades not velvety, usually 5+ times as long as wide; calyces obscurely ribbed, stellate-hairy and with long simple hairs; inflorescences axillary solitary flowers; fruits 5–6 mm diam.; mericarp spines to 1 mm; Texas. |
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17 Sida tragiifolia |
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| 11 (9) |
Leaf blade margins usually entire basally, distally dentate or serrate |
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(12) |
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Leaf blade margins crenulate-serrate to dentate to base |
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(14) |
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| 12 (11) |
Pedicels 0.4–1.2 cm; leaf blades lanceolate-elliptic to round, 1.5–5 cm. |
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2 Sida antillensis |
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Pedicels 0.5–4(–16) cm; leaf blades rhombic, subrhombic, or elliptic, 2.5–9 cm |
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(13) |
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| 13 (12) |
Petals yellow; stem hairs to 0.1 mm; pedicels (1–)3–4 cm. |
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13 Sida rhombifolia |
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Petals cream or pale yellow with reddish spot at base; stem hairs to 0.5 mm; pedicels to 2 cm. |
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15 Sida santaremensis |
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| 14 (11) |
Pedicels 8–12(–16) cm. |
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10 Sida longipes |
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Pedicels 0.5–6 cm |
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(15) |
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| 15 (14) |
Leaf blades narrowly oblong-lanceolate or elliptic to linear, 4–20 times as long as wide |
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(16) |
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Leaf blades narrowly lanceolate or elliptic to subrhombic, 2.5–10 times as long as wide |
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(17) |
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| 16 (15) |
Plants usually little-branched from base; flowers little, if at all, apically congested; petals yellow-orange. |
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5 Sida elliottii |
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Plants freely branching from base; flowers apically congested; petals yellow-orange to reddish, sometimes drying lavender. |
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11 Sida neomexicana |
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| 17 (15) |
Pedicels 2–6 cm, sometimes equaling subtending leaf; mericarps 8–10. |
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8 Sida lindheimeri |
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Pedicels 0.5–3 cm, shorter than subtending leaf; mericarps 9–12 |
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(18) |
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| 18 (17) |
Pedicels unarticulated; stipules subequal to corresponding petioles. |
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9 Sida littoralis |
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Pedicels articulated; stipules 2 times length of corresponding petioles. |
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14 Sida rubromarginata |
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List of lower taxa
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