Plants green to tan. Stem central strand usually present, sclerodermis and hyalodermis commonly absent. Leaves usually ovate or broadly ligulate to spathulate, apex usually rounded or broadly acute, margins recurved basally, distal laminal cells if 2-stratose stacked directly over one another, walls smooth or papillose, KOH yellow or red, occasionally orange, occasionally yellow distally and red basally; costa with or without a differentiated abaxial epidermis; gemmae largely absent.

Genera 21 (15 in the flora): worldwide.