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FNA | Family List | FNA Vol. 20 | Asteraceae | Solidago

52. Solidago rugosa Miller, Gard. Dict. ed. 8. Solidago no. 25. 1768.

Rough-stemmed or wrinkle-leaf goldenrod , verge d’or rugueuse

Aster rugosus (Miller) Kuntze

Plants 30–200 cm; rhizomes long-creeping, forming clones. Stems 1–50+, erect, glabrous or densely hispid to strigose. Leaves: basal withering by flowering; proximal cauline usually withering by flowering, sessile, blades elliptic to lanceolate, 68–104 × 20–25 mm, margins sharply serrate, apices acute to attenuate, abaxial faces usually hispido-strigose (at least on main nerves), nerves sometimes prominent, abaxial glabrate; mid to distal cauline sessile, blades lanceolate, elliptic or ovate, (15–)40–75(–90) × (6–)12–22(–32) mm, largest at midstem, somewhat reduced to much reduced distally, margins coarsely to finely serrate, ciliate, indument similar to proximal or denser. Heads 50–1500 , secund, in secund-pyramidal paniculiform arrays 7–36(–50) × 9–26 cm, compact to lax, branches divergent and recurved, longest 0.8–34 cm, leafy-bracteate. Peduncles 1–1.8 mm, sparsely to densely hispido-strigillose; bracteoles linear- lanceolate to ovate. Involucres narrowly campanulate, (2–)2.5–3.5(–4.5) mm. Phyllaries in 3–4 series, strongly unequal, lanceolate to linear-lanceolate, acute to obtuse. Ray florets (4–)6–8(–12); laminae (0.9–)1–1.6(–2.3) × 0.4–0.7 mm. Disc florets (2–)4–6(–8); corollas 2–3.5(–4.5) mm, lobes (0.5–)0.7–1(–1.3) mm. Cypselae (narrowly obconic) 0.9–1.5 mm, moderately strigillose; pappi 1.8–2.5 mm.

Subspecies 2 (2 in the flora): e North America.

Solidago rugosa is highly variable in size, array shape, and hairiness. It is similar to members of the S. canadensis complex; it differs in not having 3-nerved leaves. The species is divided into two subspecies and five varieties that can be difficult to distinguish.

1 Leaves relatively thin, not very rugose, usually sharply toothed, apices acuminate, glabrous or relatively soft-hairy; ray florets (4–)6–11(–13); northern (subsp. rugosa)   (2)
+ Leaves relatively thick and firm, strongly rugose-nerved, usually blunt-toothed to subentire, apices often acute, relatively short and stiff hairy; ray florets 4–9; mostly se United States (subsp. aspera)   (3)
2 (1) Stems and leaves hairy; e Canada and ne United States s to Virginia   52a Solidago rugosa var. rugosa
+ Stems and leaves glabrous; coastal cedar bogs and swamps   52b Solidago rugosa var. sphagnophila
3 (1) Arrays narrow, proximal branches not much exceeding subtending leaves; herbage sparsely hairy; mid to higher elevations in mountains   52e Solidago rugosa var. cronquistiana
+ Arrays wide, usually with elongate proximal branches greatly exceeding subtending leaves; herbage moderately to densely hairy; lower elevations in mountains, piedmont, and coastal plain   (4)
4 (3) Distal cauline leaves lanceolate to elliptic, not much reduced distally; much of range of sub-species   52c Solidago rugosa var. aspera
+ Distal cauline leaves ovate, much reduced distally; outer coastal plain   52d Solidago rugosa var. celtidifolia


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