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FNA | Family List | FNA Vol. 28 | Orthotrichaceae | Orthotrichum

33. Orthotrichum pycnophyllum Schimper in J. K. A. Müller, Syn. Musc. Frond. 1: 709. 1849.

Orthotrichum lozanoi Cardot; O. lozanoi var. lutescens Cardot; O. pycnophyllum var. lutescens (Cardot) Grout; O. recurvans Müller Hal.

Plants 0.8-4 cm. Stem leaves loosely erect-spreading when dry, lanceolate, 2.5-4 mm; margins revolute to just below apex, often wavy and slightly notched near apex; apex gradually acuminate to long acuminate-cuspidate; basal laminal cells elongate, walls thick, nodose; distal cells 7-13 µm, 1-stratose, papillae 1 or 2 per cell, conic, size moderate. Specialized asexual reproduction absent. Sexual condition gonioautoicous. Seta 1-1.5 mm. Capsule immersed to fully exserted, usually 1/2 -1/4 emergent, oblong-ovate to cylindric, 1.5-2 mm, smooth to slightly 8-ribbed; stomata superficial; peristome double; prostome absent; exostome teeth 16, , recurved, , obscurely and densely papillose; endostome segments 16, separate to base, well developed, , of 2 rows of cells, , coarsely papillose. Calyptra conic-oblong, smooth, moderately hairy, hairs smooth. Spores 17-29 µm.

Branches and trunks of trees; moderate to high elevations (500-2000 m); Ariz., N.Mex., Tex.; Mexico; Central America (Guatemala, Panama); South America.

Distinguishing features of Orthotrichum pycnophyllum are the 16 exostome teeth that are not connate at the base to form a membrane as in O. keeverae, 16 endostome segments, and a slightly ribbed capsule that is usually 3/4 emergent.


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