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FNA | Family List | FNA Vol. 27 | Pottiaceae | Tortula

14. Tortula modica R. H. Zander, Bull. Buffalo Soc. Nat. Sci. 32: 226. 1993.

Gymnostomum intermedium Turner, Muscol. Hibern. Spic., 7, plate 1, figs. a-c. 1801, not Tortula intermedia (Bridel) De Notaris 1838; Pottia intermedia (Turner) Fürnrohr; P. truncata var. major (F. Weber & D. Mohr) Bruch & Schimper

Leaves obovate to spatulate, apex broadly acute or occasionally rounded, short-awned, margins weakly recurved proximally or occasionally plane, weakly bordered distally with 2-4 rows of slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2-3 cells across the convex adaxial surface; distal laminal cells irregularly hexagonal or sometimes rectangular, width 20-25(-29) µm wide, 1(-2):1, smooth or 1-papillose (best seen in section). Sexual condition autoicous. Sporophytes exerted. Seta 0.4-0.6 cm. Capsule stegocarpic, not systylius, short-cylindric, erect and nearly straight, urn 1-1.5 mm; peristome absent or rudimentary; operculum ca. 0.5 mm. Spores 30-35 µm, spheric, densely papillose.

Capsules mature fall-spring. Calcareous soil, old fields, pastures, rocky areas; low and moderate elevations; Ont.; N.Y., Pa.; Europe; Asia; n Africa.

A report of Tortula modica from Mexico was excluded by R. H. Zander (1993). The distribution and characters of this uncommon species were discussed by H. A. Crum and L. E. Anderson (1981), P. M. Eckel (1987), and C. Williams (1966b). Essentially, Tortula modica is a high polyploid (n = 52) taxon distinguished from T. truncata by the larger habit, leaf margins usually recurved proximally, capsules slightly longer, short-cylindric, not tapering to the base, and peristome sometimes present though rudimentary.


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