Physalis arenicola var. ciliosa (Rydberg) Waterfall; P. ciliosa Rydberg

Herbs perennial, rhizomatous, rhizomes deeply buried, slen­der, typically also with shal­lowly buried, slender rhizomes, glabrous to villous, hairs simple, antrorse, to 1 mm, sometimes also with simple, jointed, divergent hairs, 1–2 mm, sometimes glandular. Stems erect, few-branched, 0.5–3 dm. Leaves petiolate; petiole 1/4–2/3 blade; blade ovate to suborbiculate, 1.5–6(–6.5) × 1–5 cm, base truncate to cordate, margins entire or coarsely, irregularly dentate with few teeth. Pedicels (8–)11–17(–25) mm, 15–30(–35) mm in fruit. Flowers: calyx 6–12 mm, villous, lobes 2–5 mm; corolla yellow with 5 pale reddish-brown smudges or not, campanulate-rotate, 10–17 mm; anthers yellow, not twisted after dehiscence, 2.5–4 mm. Fruiting calyces loosely enclosing berry, 10-ribbed, 20–35 × 15–25 mm. 2n = 24.

Flowering year-round in areas without frost. Sandy soil, pine-oak woods, hammocks, fields, pastures, road­sides; 0–50 m; Ala., Fla., Ga., Miss.

Physalis arenicola is found throughout Florida; only a few records exist from the other states in its range.