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17. Drypetes Vahl, Eclogae Amer.  3: 49.  1810.  
核果木属 he guo mu shu 
 
Authors: Bingtao Li & Michael G. Gilbert
 
 
 
Cyclostemon Blume. 
Trees or shrubs, dioecious; indumentum of simple hairs, often absent. Leaves alternate; stipules 2; petioles short; leaf blade simple, base often asymmetrical, margin entire or serrate, venation pinnate. Inflorescences axillary, sometimes grouped into glomerules, racemes, or panicles. Male flowers pedicellate or sessile; sepals 4-6, distinct, usually unequal, imbricate; petals absent; stamens 1-25, 1 to several whorls surrounding disk or outer ones inserted at margin or sinus of disk, inner ones inserted on disk; filaments free; anthers bilocular, usually introrse, rarely extrorse, longitudinally dehiscent; disk flattened or concave, margin shallowly or deeply fid; rudimentary ovary very small or absent. Female flowers solitary, axillary, sometimes on older branches; sepals as in male; petals absent; disk annular; ovary 1- or 2(or 3)-loculed; ovules 2 per locule; style short, stigmatiform; stigmas 1 or 2(or 3), usually dilated, peltate or reniform. Fruit a drupe, subglobose or ovoid, 1- or 2(or 3)-celled, seeds 1 per locule; exocarp leathery or nearly so; mesocarp fleshy or woody; endocarp woody, papery, or crustaceous. Seeds without caruncle; endosperm fleshy; cotyledons larger and flattened. 
About 200 species: tropical and subtropical Africa, America, and Asia; 12 species (two endemic) in China. 
 
 
 
The timber from several species is valued. 
 
 
 
 
                        
                         
		
                          
                          
			
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                                  Key to material with male flowers (not known for D. integrifolia, D. littoralis, D. longistipitata, and D. obtusa) 
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  | 1 | 
  Male pedicels 13-18 mm. | 
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  7 D. cumingii | 
 
  | + | 
  Male pedicels to 5 mm | 
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  (2) | 
 
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  | 2 (1) | 
  Stamens 4-8, inserted in sinuses between disk lobes; flowers in elongated inflorescences, branched or unbranched | 
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  (3) | 
 
  | + | 
  Stamens 10-25, disk annular; flowers in sessile clusters | 
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  (4) | 
 
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  | 3 (2) | 
  Leaf blade margin toothed, reticulate veins distant; filaments short and slightly flattened; shrub to 4 m. | 
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  3 D. arcuatinervia | 
 
  | + | 
  Leaf blade margin entire, reticulate veins dense and prominent; filaments terete; trees to 15 m. | 
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  11 D. indica | 
 
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  | 4 (2) | 
  Stamens 18-25, filaments flattened, wider at base | 
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  (5) | 
 
  | + | 
  Stamens 10-15, filaments slender, terete | 
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  (6) | 
 
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  | 5 (4) | 
  Petiole 8-10 mm; leaf apex obtuse, margin entire. | 
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  9 D. hainanensis | 
 
  | + | 
  Petiole 3-6 mm; leaf apex acute, margin obtusely serrate. | 
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  1 D. perreticulata | 
 
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  | 6 (4) | 
  Branchlets pubescent; leaf blade 4.5-6 cm, lateral veins 7-9 pairs. | 
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  6 D. congestiflora | 
 
  | + | 
  Branchlets glabrous; leaf blade 6-11 cm, lateral veins 8-14 pairs | 
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  (7) | 
 
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  | 7 (6) | 
  Flowers 1-3 per cluster, with cup-shaped rudimentary ovary. | 
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  12 D. salicifolia | 
 
  | + | 
  Flowers 3-5 per cluster, without rudimentary ovary. | 
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  2 D. hoaensis | 
 
 
 
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                                List of Keys
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                          List of lower taxa 
 
                        
                          
                        
	
                        
                      
 
                      
	 
                      
                         
		
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