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17. Drypetes Vahl, Eclogae Amer. 3: 49. 1810.
核果木属 he guo mu shu
Authors: Bingtao Li & Michael G. Gilbert
Cyclostemon Blume.
Trees or shrubs, dioecious; indumentum of simple hairs, often absent. Leaves alternate; stipules 2; petioles short; leaf blade simple, base often asymmetrical, margin entire or serrate, venation pinnate. Inflorescences axillary, sometimes grouped into glomerules, racemes, or panicles. Male flowers pedicellate or sessile; sepals 4-6, distinct, usually unequal, imbricate; petals absent; stamens 1-25, 1 to several whorls surrounding disk or outer ones inserted at margin or sinus of disk, inner ones inserted on disk; filaments free; anthers bilocular, usually introrse, rarely extrorse, longitudinally dehiscent; disk flattened or concave, margin shallowly or deeply fid; rudimentary ovary very small or absent. Female flowers solitary, axillary, sometimes on older branches; sepals as in male; petals absent; disk annular; ovary 1- or 2(or 3)-loculed; ovules 2 per locule; style short, stigmatiform; stigmas 1 or 2(or 3), usually dilated, peltate or reniform. Fruit a drupe, subglobose or ovoid, 1- or 2(or 3)-celled, seeds 1 per locule; exocarp leathery or nearly so; mesocarp fleshy or woody; endocarp woody, papery, or crustaceous. Seeds without caruncle; endosperm fleshy; cotyledons larger and flattened.
About 200 species: tropical and subtropical Africa, America, and Asia; 12 species (two endemic) in China.
The timber from several species is valued.
Key to fruiting material
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1 |
Ovary and drupe 1-locular |
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(2) |
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Ovary and drupe 2(or 3)-locular |
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(6) |
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2 (1) |
Fruits glabrous |
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(3) |
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Fruits pubescent |
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(4) |
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3 (2) |
Branchlets and petioles puberulent when young; lateral veins 6-8 pairs; stipules linear, persistent; stamens ca. 25; fruits 1.8-2.5 cm. |
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1 D. perreticulata |
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Branchlets and petioles glabrous when young; lateral veins 9-13 pairs; stipules small, caducous; stamens 12-15; fruits 1-2 cm. |
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2 D. hoaensis |
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4 (2) |
Leaf blade acuminate at apex, margin sparsely obtuse-serrate, lateral veins prominent. |
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3 D. arcuatinervia |
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Leaf blade shortly acuminate at apex, margin entire or sometimes obscurely obtuse-serrate at upper part, lateral veins obscure |
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(5) |
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5 (4) |
Trees; branchlets and petioles pubescent when young; fruits angular and rugose. |
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4 D. obtusa |
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Shrubs; branchlets and petioles glabrous; fruits rounded and smooth. |
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5 D. integrifolia |
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6 (1) |
Leaf margins serrate |
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(7) |
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Leaf margins entire |
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(8) |
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7 (6) |
Branchlets and petioles pubescent; pedicels 2-4 mm; stamens 13-15; fruits globose, glabrous. |
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6 D. congestiflora |
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Branchlets and petioles glabrous; pedicels 13-18 mm; stamens 10; fruits oblong or ellipsoid, pubescent. |
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7 D. cumingii |
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8 (6) |
Fruits ellipsoid-ovoid; leaf blade slightly falcate. |
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8 D. littoralis |
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Fruits globose or subglobose; leaf blade not falcate |
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(9) |
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9 (8) |
Leaf apex obtuse |
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(10) |
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Leaf apex caudate-acuminate or acuminate |
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(11) |
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10 (9) |
Fruiting pedicels stout, ca. 0.5 cm; leaf blade with 8-10 pairs of lateral veins; fruits 2-2.5 cm in diam.. |
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9 D. hainanensis |
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Fruiting pedicels slender, 1.5-2 cm; leaf blade with 4 or 5 pairs of lateral veins; fruits ca. 1 cm in diam.. |
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10 D. longistipitata |
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11 (9) |
Female pedicels to 2 cm, 2.5-4 cm in fruit; leaf blade elliptic-oblong to lanceolate; stamens 4-8. |
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11 D. indica |
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Female pedicels 0.2-0.4 cm in fruit; leaf blade linear-oblong; stamens 12. |
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12 D. salicifolia |
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List of Keys
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List of lower taxa
Related Links (opens in a new window) |
Treatments in Other Floras @ www.efloras.org
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