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FOC | Family List | FOC Vol. 9 | Rosaceae

43. AGRIMONIA Linnaeus, Sp. Pl. 1: 448. 1753.

龙芽草属 long ya cao shu

Li Chaoluan (Li Chao-luang); Hiroshi Ikeda, Hideaki Ohba

Herbs perennial, tall, with creeping rhizome. Leaves stipulate, imparipinnate. Flowers usually in terminal, spikelike racemes, bisexual, rather small. Hypanthium turbinate, sulcate, with hooked prickles or 5 teeth below limb abaxially, constricted at throat. Sepals 5, imbricate, persistent. Petals 5, larger than sepals. Disk lining hypanthium, margin thickened, annular, glandular. Stamens 5–15 or more, inserted at mouth of hypanthium, 1-seriate. Carpels usually 2, included in hypanthium, sessile; ovule pendulous; style terminal, exserted, filiform; stigma dilated. Achenes 1 or 2, included in closed hypanthium, distally with hooked prickles. Seed pendulous; testa membranous. x = 7.

About ten species: N temperate zone, alpine regions of tropics; four species in China.

1 Flowers 12–13 mm in diam.; fruiting hypanthium 8–10 × ca. 5 mm including prickles, outer prickles recurved, innermost ones spreading.   4 A. eupatoria subsp. asiatica
+ Flowers 4–10 mm in diam.; fruiting hypanthium 5–8 × 2–4 mm including prickles, prickles spreading, erect, or connivent.   (2)
2 (1) Stipules flabellate or broadly ovate, margin obtusely dentate; stamens 17–24.   3 A. coreana
+ Stipules falcate or semiorbicular, margin acutely serrate; stamens 5–15.   (3)
3 (2) Flowers 6–9 mm in diam.; fruiting hypanthium 7–8 × 3–4 mm including prickles, prickles erect when young, connivent when old; leaflets obovate to obovate-lanceolate, abaxially appressed pubescent on veins.   1 A. pilosa
+ Flowers 4–5 mm in diam.; fruiting hypanthium 4–5 × 2–2.5 mm including prickles, prickles spreading; leaflets rhombic-elliptic or elliptic, usually widest at middle, abaxially sparsely spreading hirsute on veins.   2 A. nipponica var. occidentalis

Lower Taxa


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