1 |
Involucres straw-colored, glossy, 2-4 mm in diam.; phyllaries with colorless scarious margin |
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(2) |
+ |
Involucres not straw-colored, not glossy, 4-10 mm in diam.; phyllaries with dark brown or purple scarious margin |
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(15) |
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2 (1) |
Capitula 5-14, often regularly fascicled-corymbose at apices of branches; involucres cylindric or narrowly campanulate; phyllaries rigidly herbaceous, apex acute |
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(3) |
+ |
Capitula many, corymbose or compound-corymbose at apices of stem and branches; involucres campanulate; phyllaries herbaceous, apex obtuse or rounded |
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(4) |
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3 (2) |
Leaf blade adaxially green, 2-pinnatisect. |
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23 A. parviflora |
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Leaf blade gray-green, 3-parted or -lobed. |
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24 A. trifida |
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4 (2) |
Perennial herbs |
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(5) |
+ |
Small shrubs or subshrubs |
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(7) |
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5 (4) |
Involucres 6-7 mm in diam.; outer phyllaries narrowly elliptic, apices with rounded waxy appendage. |
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33 A. pallasiana |
+ |
Involucres 2.5-4 mm in diam.; outer phyllaries ovate or lanceolate, apices without rounded waxy appendage |
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(6) |
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6 (5) |
Stem weakly branched at base, often solitary, almost herbaceous; middle and lower stem leaf blades usually bipinnate with 5-7 lobes. |
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34 A. fastigiata |
+ |
Stem strongly branched at base, woody for much of length; middle and lower stem leaf blades usually twice ternately divided. |
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35 A. gracilis |
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7 (4) |
Young stems glabrous; leaf blade abaxially green or pale green, glabrous or nearly so. |
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32 A. nematoloba |
+ |
Young stems hairy; leaf blade abaxially gray-white, appressed pubescent |
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(8) |
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8 (7) |
Leaf blade shallowly toothed to subentire, only occasionally lobed |
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(9) |
+ |
Leaf blade pinnatisect, pinnatipartite, 2-pinnatisect, palmate-pinnately 3-5-divided, or twice ternately divided |
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(10) |
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9 (8) |
Leaf blade adaxially green or gray-green, glabrous or very sparsely pubescent; disk florets ca. 2.5 mm. |
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25 A. potaninii |
+ |
Leaf blade adaxially densely gray-white tomentose; disk florets ca. 4 mm. |
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26 A. amphisericea |
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10 (8) |
Leaf blade pinnatisect or pinnatipartite |
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(11) |
+ |
Leaf blade 2-pinnatisect, palmate-pinnately 3-5-divided, bipalmatisect, or twice ternately divided |
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(12) |
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11 (10) |
Leaf blade pinnatisect, adaxially gray-white, densely pubescent. |
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27 A. truncata |
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Leaf blade pinnatipartite, adaxially green, glabrous. |
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28 A. hypoleuca |
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12 (10) |
Leaf blade 2-pinnatisect. |
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29 A. achilleoides |
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Leaf blade bipalmatisect, palmately-pinnately 3-5-divided, or twice ternately divided |
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(13) |
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13 (12) |
Leaf blade twice ternately divided. |
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35 A. gracilis |
+ |
Leaf blade bipalmately 3-5-sect or bipalmate, pinnately 3-5-sect |
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(14) |
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14 (13) |
Leaves with divided pseudostipules. |
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30 A. nitida |
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Leaves with entire pseudostipules. |
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31 A. fruticulosa |
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15 (1) |
Leaf blade undivided, 3-lobed, or simply pinnatisect, or pinnatipartite |
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(16) |
+ |
Leaf blade more complexly divided: 2-pinnatisect, bipalmatisect, palmate-pinnately 3-7-divided, tripinnatisect, or tripinnate, pinnately divided |
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(21) |
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16 (15) |
Leaf blade undivided, margin entire, serrate, or incised-serrate |
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(17) |
+ |
Leaf blade 3-lobed or pinnately divided |
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(19) |
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17 (16) |
Leaf blade linear, margin entire. |
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1 A. salicifolia |
+ |
Leaf blade narrowly elliptic, lanceolate, or obovate-oblong, or broadly oblanceolate, rarely linear, margin serrate or incised-serrate |
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(18) |
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18 (17) |
Subshrubs; leaves 5-10 × 0.3-4 cm, without pseudostipules, abaxially densely and thickly tomentose. |
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2 A. quercifolia |
+ |
Perennial herbs; leaves 3-4.5 × 1-2 cm, with divided pseudostipules, abaxially densely pubescent. |
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3 A. latifolia |
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19 (16) |
Leaf blade 3-lobed. |
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6 A. alabasica |
+ |
Leaf blade pinnately divided |
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(20) |
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20 (19) |
Leaf blade with 1 or 2 pairs of lateral lobes; lobes linear. |
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4 A. variifolia |
+ |
Leaf blade with 3 or 4 pairs of lateral lobes; lobes narrowly elliptic, lanceolate, falcate, or obliquely triangular. |
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5 A. ramosa |
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21 (15) |
Small subshrubs with thick, woody rootstocks; stems 4-25 cm |
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(22) |
+ |
Erect or diffuse perennial herbs, with slender fibrous roots; stems often more than 25 cm |
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(25) |
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22 (21) |
Capitula solitary; involucres 7-10 mm in diam. |
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10 A. scharnhorstii |
+ |
Capitula in panicles; involucres 4-7(-8) mm in diam |
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(23) |
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23 (22) |
Phyllary margin dark purple; corolla apex purple-red. |
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8 A. purpurea |
+ |
Phyllary margin deep brown; florets uniformly yellow |
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(24) |
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24 (23) |
Leaf blade elliptic or obliquely elliptic in outline, 1-2 cm, both surfaces densely tomentose; involucre 4-6 mm in diam. |
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7 A. tibetica |
+ |
Leaf blade suborbicular in outline, 0.3-0.5 cm, both surfaces densely pubescent; involucre 5-7 mm in diam. |
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9 A. trilobata |
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25 (21) |
Leaf blade tripinnatisect, subtripinnatisect, or palmate, tripinnately divided |
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(26) |
+ |
Leaf blade 2-pinnatisect, bipalmatisect, or subpalmately 3-5-divided |
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(29) |
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26 (25) |
Whole plant densely and thickly tomentose; phyllaries inconspicuous. |
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19 A. sericea |
+ |
Whole plant pubescent or densely so; phyllaries conspicuous |
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(27) |
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27 (26) |
Leaf blade 3-3.5 × 2-3 cm, adaxially green or pale green, glabrous or nearly so, ultimate segments lanceolate or elliptic. |
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20 A. tripinnatisecta |
+ |
Leaf blade 1-2 × 1-1.5 cm, adaxially gray-white, densely pubescent, ultimate segments linear |
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(28) |
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28 (27) |
Stems thinly silvery gray tomentose; leaf blade ca. 2 × 1.5 cm. |
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21 A. elegantula |
+ |
Stems densely pubescent; leaf blade ca. 1 × 1 cm. |
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22 A. nubigena |
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29 (25) |
Leaf blade bipalmatisect or palmately 3-5-divided |
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(30) |
+ |
Leaf blade 2-pinnatisect |
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(31) |
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30 (29) |
Leaf blade adaxially green, subglabrous; involucres 4-5 mm in diam. |
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17 A. breviloba |
+ |
Leaf blade adaxially densely gray-white or white pubescent; involucres 6-10 mm in diam. |
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18 A. khartensis |
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31 (29) |
Phyllary margin ± distinctly 2-colored: dark brown with a narrow but distinct colorless margin. |
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16 A. tenuifolia |
+ |
Phyllary margin uniformly brown or dark brown scarious |
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(32) |
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32 (31) |
Primary lateral segments of leaf blade pinnatilobed, pinnatifid, or pinnatipartite |
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(33) |
+ |
Primary lateral segments of leaf blade or leaf blade pinnatisect |
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(34) |
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33 (32) |
Primary lateral segments of leaf blade 1- or 2-paired, if 2-paired then ca. 1 cm apart; ultimate lobes narrowly elliptic or falcate. |
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14 A. remotipinna |
+ |
Primary lateral segments of leaf blade 2-4-paired, ca. 5 mm at intervals; ultimate lobes elliptic, lanceolate, or obliquely triangular. |
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15 A. myriantha |
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34 (32) |
Leaf blade adaxially gray-white, densely villous or pubescent; involucres 5-6 mm in diam.; corolla of marginal female florets with truncate apex. |
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11 A. brachyantha |
+ |
Leaf blade adaxially green, subglabrous; involucres 2.5-5 mm in diam.; corolla of marginal female florets with toothed apex |
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(35) |
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35 (34) |
Ultimate segments of leaf blade linear-lanceolate or narrowly elliptic; involucres ca. 2.5 mm in diam. |
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12 A. przewalskii |
+ |
Ultimate segments of leaf blade linear; involucres 4-5 mm in diam. |
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13 A. adenantha |