| 1 |
Leaf sheaths pubescent or spinulose. |
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(2) |
| + |
Leaf sheaths smooth (occasionally slightly pubescent in A. biserrata). |
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(6) |
| |
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|
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| 2 (1) |
Leaf rachis densely pubescent. |
|
1 Angelica duclouxii |
| + |
Leaf rachis glabrous. |
|
(3) |
| |
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| 3 (2) |
Leaves glabrous. |
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(4) |
| + |
Leaves hispid or setulose along nerves. |
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(5) |
| |
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|
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| 4 (3) |
Bracts 5–9; vittae absent on commissure. |
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2 Angelica apaensis |
| + |
Bracts absent or 1; vittae 2 on commissure. |
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3 Angelica anomala |
| |
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| 5 (3) |
Leaf sheaths spinulose; fruit ellipsoid to narrow-ellipsoid; vittae 4 on commissure. |
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4 Angelica valida |
| + |
Leaf sheaths pubescent; fruit suborbicular to ellipsoid; vittae 2 on commissure. |
|
5 Angelica setchuenensis |
| |
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|
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| 6 (1) |
Rachis and petiolules geniculate. |
|
(7) |
| + |
Rachis and petiolules not geniculate. |
|
(10) |
| |
|
|
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| 7 (6) |
Ovary puberulent or hispidulous. |
|
(8) |
| + |
Ovary glabrous. |
|
(9) |
| |
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|
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| 8 (7) |
Bracteoles absent. |
|
6 Angelica fargesii |
| + |
Bracteoles many, linear. |
|
7 Angelica genuflexa |
| |
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|
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| 9 (7) |
Bracteoles white-scarious-margined; fruit narrow-oblong, 6–7 × 3–3.5 mm. |
|
8 Angelica tianmuensis |
| + |
Bracteoles not white-scarious-margined, fruit oblong-ellipsoid, 6–7 × 3–5 mm. |
|
9 Angelica polymorpha |
| |
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|
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| 10 (6) |
Basal and lower cauline leaves 1–4-pinnate. |
|
(11) |
| + |
Basal and lower cauline leaves 1–3-ternate or 1–3-ternate-pinnate. |
|
(21) |
| |
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|
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| 11 (10) |
Rays 7–20. |
|
(12) |
| + |
Rays 20–50. |
|
(14) |
| |
|
|
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| 12 (11) |
Apex of leaves obtuse. |
|
10 Angelica nitida |
| + |
Apex of leaves acute to long-acuminate. |
|
(13) |
| |
|
|
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| 13 (12) |
Bracteoles lanceolate, apex long-aristate. |
|
11 Angelica dielsii |
| + |
Bracteoles small, subulate. |
|
12 Angelica omeiensis |
| |
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|
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| 14 (11) |
Leaves 2–4-pinnate. |
|
(15) |
| + |
Leaves 1–2-pinnate. |
|
(17) |
| |
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|
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| 15 (14) |
Leaflet margin ciliate, apex long-caudate-acuminate. |
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18 Angelica longipes |
| + |
Leaflet margin not ciliate, apex acute. |
|
(16) |
| |
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| 16 (15) |
Petals white; calyx teeth obsolete. |
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19 Angelica sylvestris |
| + |
Petals greenish; calyx teeth conspicuous, triangular-ovate. |
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20 Angelica dailingensis |
| |
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| 17 (14) |
Leaves pinnate. |
|
|
| + |
Leaves 2-pinnate. |
|
(19) |
| |
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|
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| 18 |
Pedicels 10–25 mm (Yunnan). |
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13 Angelica longipedicellata |
| + |
Pedicels 4–7 mm (Taiwan). |
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14 Angelica tarokoensis |
| |
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| 19 (17) |
Leaflets glabrous. |
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15 Angelica songpanensis |
| + |
Leaflets pubescent along nerves. |
|
(20) |
| |
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|
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| 20 (19) |
Bract and bracteole margin ciliate; rays densely pubescent. |
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16 Angelica pseudoselinum |
| + |
Bract and bracteole margin not ciliate; rays subglabrous. |
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17 Angelica morrisonicola |
| |
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| 21 (10) |
Leaves 1–3-ternate. |
|
(22) |
| + |
Leaves 1–3-ternate-pinnate. |
|
(23) |
| |
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| 22 (21) |
Apex of leaves acuminate; vittae 2–3 in each furrow, 4 on commissure (Gansu, Shaanxi). |
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21 Angelica tsinlingensis |
| + |
Apex of leaves obtuse-rounded or acute; vittae 1 in each furrow, 2 on commissure (Xinjiang). |
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22 Angelica ternata |
| |
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| 23 (21) |
Petals hirsute abaxially; ovary hispid. |
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23 Angelica hirsutiflora |
| + |
Petals and ovary glabrous (ovary pubescent in A. dahurica var. formosana). |
|
(24) |
| |
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| 24 (23) |
Calyx teeth conspicuous, triangular-ovate to subulate. |
|
(25) |
| + |
Calyx teeth obsolete. |
|
(28) |
| |
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| 25 (24) |
Leaves ternate to ternate-pinnate; secondary ribs of fruit 2, slightly prominent. |
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24 Angelica oncosepala |
| + |
Leaves 1–2-ternate-pinnate; secondary ribs of fruit not developed. |
|
(26) |
| |
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| 26 (25) |
Bracteoles pinnate. |
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25 Angelica pinnatiloba |
| + |
Bracteoles not pinnate. |
|
(27) |
| |
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| 27 (26) |
Leaflets decurrent on petioles, hispidulous adaxially. |
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26 Angelica decursiva |
| + |
Leaflets not decurrent, glabrous. |
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28 Angelica kangdingensis |
| |
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| 28 (24) |
Bract and bracteole margin ciliate. |
|
(29) |
| + |
Bract and bracteole margin not ciliate. |
|
(32) |
| |
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| 29 (28) |
Stem glabrous. |
|
(30) |
| + |
Stem pubescent or hispidulous. |
|
(31) |
| |
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| 30 (29) |
Fruit narrow-oblong, 5–9 × 2.5–4 mm; vittae 3–6 on commissure. |
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29 Angelica longicaudata |
| + |
Fruit suborbicular, 4–6 × 3–5 mm; vittae 2 on commissure. |
|
30 Angelica laxifoliata |
| |
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| 31 (29) |
Leaflet margin serrulate and ciliate, rays 40–60. |
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31 Angelica maowenensis |
| + |
Leaflet margin irregularly biserrate; rays 10–25. |
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32 Angelica biserrata |
| |
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| 32 (28) |
Leaflet base decurrent, rachis conspicuously winged. |
|
(33) |
| + |
Leaflets base not decurrent (A. dahurica slightly decurrent), rachis not winged. |
|
(34) |
| |
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|
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| 33 (32) |
Bracts absent; petals white. |
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33 Angelica cartilaginomarginata |
| + |
Bracts 2; petals dark purple-red. |
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27 Angelica gigas |
| |
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| 34 (32) |
Stem and leaves glabrous. |
|
(35) |
| + |
Stem and leaves usually hairy. |
|
(38) |
| |
|
|
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| 35 (34) |
Bracts absent; fruit narrow-oblong. |
|
34 Angelica acutiloba |
| + |
Bracts developed; fruit ellipsoid to suborbicular. |
|
(36) |
| |
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|
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| 36 (35) |
Rays 17–30, unequal; fruit 5–7 mm long. |
|
35 Angelica paeoniifolia |
| + |
Rays 10–20, subequal; fruit 7–12 mm long. |
|
(37) |
| |
|
|
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| 37 (36) |
Proximal pinnules of leaves 3-foliolose; dorsal ribs of fruit equally winged (Xizang). |
|
36 Angelica glauca |
| + |
Proximal pinnules of leaves not 3-foliolose; dorsal ribs of fruit unequally winged (Xinjiang). |
|
37 Angelica multicaulis |
| |
|
|
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| 38 (34) |
Rachis, peduncles, rays and pedicels all densely hispidulous. |
|
38 Angelica amurensis |
| + |
Rachis, peduncles, rays and pedicels partly hairy or glabrous. |
|
(39) |
| |
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|
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| 39 (38) |
Leaves 1–2-ternate-pinnate; bracteoles entire to 2–3-lobed. |
|
39 Angelica balangshanensis |
| + |
Leaves 2–3-ternate-pinnate; bracteoles entire. |
|
(40) |
| |
|
|
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| 40 (39) |
Leaflets 2–3.5 × 0.8–2.5 cm. |
|
(41) |
| + |
Leaflets 5–15 × 2–10 cm. |
|
(42) |
| |
|
|
|
| 41 (40) |
Bracteoles pubescent; lateral ribs of fruit narrow-winged, wings narrower than the body. |
|
40 Angelica morii |
| + |
Bracteoles glabrous; lateral ribs of fruit broad-winged, wings wider than the body. |
|
41 Angelica sinensis |
| |
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|
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| 42 (40) |
Bracteoles absent; petals dark purple-red. |
|
42 Angelica megaphylla |
| + |
Bracteoles several; petals usually white. |
|
(43) |
| |
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|
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| 43 (42) |
Leaflet margin inconspicuously serrulate; bracteole apex long-aristate. |
|
43 Angelica likiangensis |
| + |
Leaflet margin coarsely cuspidate-serrate or biserrate; bracteole apex not aristate. |
|
(44) |
| |
|
|
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| 44 (43) |
Leaflet margin ciliate; rays 16–18. |
|
44 Angelica cincta |
| + |
Leaflet margin not ciliate; rays 18–40. |
|
45 Angelica dahurica |
