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5. Arthromeris (T. Moore) J. Smith, Hist. Fil. 110. 1875.

节肢蕨属 jie zhi jue shu

Authors: Shugang Lu & Peter H. Hovenkamp

Pleopeltis sect. Arthromeris T. Moore, Index Fil. 78. 1857.

Plants, epiphytic or terrestrial, medium-sized. Rhizome long creeping, densely scaly or covered with whitish bloom and sparse scales; scales whitish, brown, or black, pseudopeltate, lanceolate-attenuate, margin entire to shallowly toothed or ciliate. Fronds remote, monomorphic, articulate to rhizome on short scaly phyllopodia. Stipe straw-colored or brown, glabrous or pubescent. Lamina imparipinnate or simple, herbaceous or leathery, glabrous or pubescent, rarely scaly abaxially. Pinnae opposite or subopposite, rarely alternate, articulate to rachis, mostly sessile or subsessile, usually oblong-lanceolate, margin entire and cartilaginous, apex acuminate to caudate; costae and lateral veins distinct; veinlets obscure, reticulate to form irregularly shaped areoles with simple or forked included veinlets. Sori orbicular or elongated, in 1 regular row on each side of costa or scattered on abaxial surface of pinnae, naked, superficial, without paraphyses. Sporangia long stalked, annulus with 14-16 cells. Spores brown, ellipsoid, surfaces verrucate with often constricted, acuminate or echinate elements. x = 12.

About 20 species: tropical and subtropical Asia; 17 species (eight endemic) in China.


1 Sori large, solitary between adjacent lateral veins, in 1 row on either side of costa   (2)
+ Sori small, more than 2 between adjacent lateral veins, scattered on either side of costa   (6)
       
2 (1) Pinnae distinctly stalked.   1 A. tenuicauda
+ Pinnae sessile   (3)
       
3 (2) Scales brown, lanceolate, acuminate at apex   (4)
+ Scales brown, dark brown, or black, long caudate at apex   (5)
       
4 (3) Sori orbicular, solitary.   2 A. wallichiana
+ Sori elliptic or 2 adnate together.   3 A. intermedia
       
5 (3) Rhizome densely scaly, scales brown; plants terrestrial.   4 A. tatsienensis
+ Rhizome sparsely scaly and with whitish bloom, scales dark brown to black; plants epiphytic.   5 A. nigropaleacea
       
6 (1) Rhizome 10-12 mm in diam.; fronds abaxially usually with whitish bloom   (7)
+ Rhizome 3-6 mm in diam.; fronds abaxially green   (10)
       
7 (6) Pinnae sessile.   14 A. wardii
+ Pinnae stalked   (8)
       
8 (7) Pinnae opposite.   15 A. medogensis
+ Pinnae alternate   (9)
       
9 (8) Lamina abaxially with glandular hairs.   16 A. caudata
+ Lamina abaxially without glandular hairs.   17 A. salicifolia
       
10 (6) Pinna apex caudate   (11)
+ Pinna apex subacute to acuminate   (13)
       
11 (10) Pinnae 5-8 pairs; lamina abaxially glabrous or pubescent.   9 A. elegans
+ Pinnae 1-4 pairs; lamina abaxially pubescent or tomentose   (12)
       
12 (11) Rhizome sparsely scaly, surface with whitish bloom; scales brown.   6 A. himalayensis
+ Rhizome densely scaly, surface concealed; scales whitish.   7 A. tomentosa
       
13 (10) Plants terrestrial; pinnae 10-15 cm; lamina abaxially glabrous.   13 A. mairei
+ Plants epiphytic; pinnae 5-12 cm; lamina abaxially glabrous, pubescent, or tomentose   (14)
       
14 (13) Pinnae (8 or)9-16 pairs; lamina abaxially tomentose   (15)
+ Pinnae 5-8 pairs, 10-12 cm; lamina abaxially glabrous or pubescent   (16)
       
15 (14) Pinnae 9-16 pairs, 5-11 cm; lamina green, abaxially tomentose.   10 A. cyrtomioides
+ Pinnae (8 or)9 or 10(-12) pairs, 5-6 cm; lamina abaxially white tomentose.   8 A. notholaenoides
       
16 (14) Lamina 15-20 cm wide, abaxially glabrous or sparsely pubescent; sori irregularly scattered in 2 or 3 ill-defined rows.   11 A. lehmannii
+ Lamina 25-30 cm wide, abaxially densely pubescent; sori in 3-5 rows, sometimes in confluent pairs.   12 A. lungtauensis

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