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FOC | Family List | FOC Vol. 16 | Convolvulaceae

10. Calystegia R. Brown, Prodr. 483. 1810.
[nom. cons.]

打碗花属 da wan hua shu

Fang Rhui-cheng; Richard K. Brummitt

Herbs prostrate or erect to twining to several meters tall, rhizomatous [or woody at base]. Leaves subsessile to petiolate, oblong to hastate or sagittate [or rarely pedate]. Inflorescences axillary, 1-flowered [or few-flowered] cymes; bracteoles 2, sepal-like, inserted immediately below calyx, ovate and sometimes saccate, enclosing calyx [or remote from calyx and subulate or leaf-shaped], persistent. Sepals subequal, persistent. Corolla white, pink [or pale yellow], funnelform, with 5 distinct midpetaline bands, glabrous. Stamens included, equal. Pollen globose, pantoporate, not spiny. Ovary 1-loculed, 4-ovuled. Style 1, included in corolla; stigmas 2, clavate. Capsule globose, glabrous, indehiscent. Seeds 4, smooth or minutely tuberculate.

Approximately 70 intergrading subspecific taxa which can be arbitrarily combined into about 25 species: mostly temperate but sparingly extending to the tropics; six species in China.

Nearly all taxa in Calystegia intergrade geographically into neighboring taxa with the exception of the widespread coastal species, C. soldanella (Linnaeus) R. Brown. Almost half of the genus is endemic in California, U.S.A., and the exceptions noted in the generic description in brackets [ ] all refer to the Californian endemics. It is impossible to draw clearly defined specific limits, and intermediate forms are always found where two taxa approximate geographically.

1 Leaves reniform; bracteoles ca. as long as or shorter than sepals; plants of seashores   1 Calystegia soldanella
+ Leaves not reniform; bracteoles longer than sepals; plants mostly of inland habitats.   (2)
2 (1) Corolla 2--3.5 cm; leaves with abruptly spreading, usually acutely 2-angled lobes   2 Calystegia hederacea
+ Corolla 4--7 cm; leaves unlobed to distinctly lobed with lobes entire to occasionally acutely 2-angled.   (3)
3 (2) Plants trailing or suberect to climbing; leaves oblong to narrowly triangular with lobes seldom more than 1/3 length of midvein, if present at all; corolla pink.   (4)
+ Plants climbing; leaves triangular with lobes 1/3--1/2 length of midvein; corolla pink or white.   (5)
4 (3) Stems appressed pubescent to villous; petiole usually less than 1.2 cm and less than 1/4 length of leaf blade; leaves without basal lobes to shortly hastate at base   3 Calystegia pellita
+ Stems glabrous to pubescent; petiole usually more than 1.2 cm and 1/5--2/3 as long as midvein; leaves distinctly lobed at base   4 Calystegia pubescens
5 (3) Corolla pink; leaf sinus with divergent sides   5 Calystegia sepium subsp. spectabilis
+ Corolla usually white; leaf sinus with parallel sides   6 Calystegia silvatica subsp. orientalis

Lower Taxa


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