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FOC | Family List | FOC Vol. 2-3 | Athyriaceae

2. Cornopteris Nakai, Bot. Mag. (Tokyo). 44: 7. 1930.

角蕨属 jiao jue shu

Authors: He Zhaorong & Masahiro Kato

Neoathyrium Ching & Z. R. Wang.

Plants hygrophytic, evergreen or summer-green. Rhizome mostly creeping, ascending or erect, few thinly creeping, apex and bases of stipes scaly; scales brown or red-brown, lanceolate, ovate-lanceolate, or ovate, entire; fronds approximate or caespitose, occasionally distant. Stipe fleshy or herbaceous, flattened when dry, base thickened and often warty, deeply grooved on adaxial side, subglabrous; lamina variably dissected from 2-pinnatilobate to 3-pinnate or more, elliptic to ovate-deltoid, apex acuminate; pinnae lanceolate or broadly lanceolate, or lower pinnae ovate-lanceolate, sessile or shortly stalked, subspreading, asymmetrical or nearly symmetrical, apex acuminate or long acuminate; pinnules often pinnatilobate; rachis and costae broadly grooved between narrow ridges on adaxial side, fleshy hornlike processes borne in grooves at base of costae and costules; veins free, pinnate, veinlets simple, bifurcate to pinnate, not reaching lamina margin. Lamina brown-green, dark brown, or black-brown, glabrous, or rachis or costae abaxially with multicellular short nodose hairs and sparsely lanceolate brown scales, rarely with unicellular short hairs. Sori linear, elliptic, or orbicular, on veinlets, exindusiate. Spores bilateral, elliptic in polar view, semicircular or reniform in equatorial view, perispore prominent, surface with few-rugate ornamentation. x = 40.

About 16 species: mainly distributed in tropical and subtropical Asia; 12 species (six endemic) in China.

Cornopteris crenulatoserrulata (species no. 12) was assigned to the monotypic genus Neoathyrium, which was recognized in FRPS (3(2): 94. 1999).

Knapp (Ferns Fern Allies Taiwan, 442. 2011) noted that Cornopteris philippinensis M. Kato has been reported for Taiwan but that the voucher specimen (Murakami, Saiki & Chen 344) differs from Philippine material and might instead correspond to a form of C. opaca. He lists its status for Taiwan as unconfirmed.

1 Sori orbicular; lamina 2- or 3-pinnate, pinnules or secondary pinnules pinnatilobate   (2)
+ Sori elliptic or linear; lamina 1-pinnate to 2-pinnate   (5)
2 (1) Rhizome long creeping   (3)
+ Rhizome ascending to erect   (4)
3 (2) Projections absent from lateral ridges of stipe base, pinnules up to 9 ?3 cm; rachis, costae, and costules abaxially with glandular unicellular hairs.   9 C. pseudofluvialis
+ Projections present on lateral ridges of stipe base, pinnules up to 4-6 ?1.5 cm; rachis, costae, and costules abaxially with eglandular unicellular hairs.   12 C. crenulatoserrulata
4 (2) Lamina 2-pinnate, at most nearly 3-pinnate; ultimate lobes entire or shallow crenate at apex.   10 C. banajaoensis
+ Lamina 3-pinnate, ultimate pinnule usually pinnatilobate; ultimate lobes acute or serrate at apex.   11 C. badia
5 (1) Lamina 2-pinnate, pinnules pinnatilobate   (6)
+ Lamina 1-pinnate, pinnae pinnatilobate with at most basal 1 (to few) pair of free adnate pinnules   (9)
6 (5) Rhizome ascending to erect.   5 C. opaca
+ Rhizome creeping   (7)
7 (6) Pinnules of lower pinnae usually obtuse-rounded at apex, rarely pointed, pinnatilobate to pinnatifid or dentate.   6 C. decurrenti-alata
+ Pinnules of lower pinnae pointed, pinnatifid to pinnatipartite   (8)
8 (7) Pinnules of basal pinnae oblong-lanceolate, base not reduced, apex shortly acuminate or acute; pinnule lobes obtuse-crenate or subentire; lamina abaxially with unicellular short hairs, rarely also with 2- or 3-celled short nodose hairs.   7 C. christenseniana
+ Pinnules of basal pinnae ovate-lanceolate or subovate, base reduced, apex usually acuminate or long acuminate; pinnule lobes pinnatilobate or crenate; lamina abaxially with multicellular short nodose hairs.   8 C. major
9 (5) Lamina narrowly elliptic-lanceolate; all pinnae sessile   (10)
+ Lamina ovate; lower pinnae shortly stalked   (11)
10 (9) Rhizome erect; pinnae contiguous, base shallowly cordate, middle part of pinna as wide as or slightly narrower than base; rachis and costae abaxially with dense curly multicellular short nodose hairs.   1 C. approximata
+ Rhizome creeping; pinnae prominently spaced, base truncate and prominently broader; rachis and costae glabrous abaxially.   2 C. latibasis
11 (9) Rhizome creeping.   6 C. decurrenti-alata
+ Rhizome ascending or erect   (12)
12 (11) Middle and upper pinnae usually pinnatifid, rarely pinnatipartite; veinlets mostly simple, rarely forked, forked veinlets mostly supramedial.   3 C. omeiensis
+ Middle and upper pinnae pinnatipartite to pinnatisect; veinlets mostly bifurcate or pinnate, forked veinlets mostly inframedial   (13)
13 (12) Basal acroscopic lobes of upper pinnae narrower and longer than basiscopic ones.   4 C. latiloba
+ Basal acroscopic pinna lobes shorter than basiscopic ones.   5 C. opaca

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