复叶角蕨 fu ye jiao jue

Plants evergreen, tall. Rhizome ascending to erect, robust, up to 5 cm in diam. including remaining stipe bases, up to 40 cm tall, apex with adpressed, entire, brown, lanceolate scales; fronds caespitose. Fertile fronds up to 2 m; stipe brown-green, up to 1 m, up to 8 mm in diam., with sparse deciduous, entire, lanceolate scales, grooved adaxially; lamina 1-3-pinnate below apex, deltoid or ovate, nearly as long as stipe, up to 75 cm wide, apex acuminate; pinnae up to 12 pairs, slightly ascending, lower pinnae opposite, with stalk up to 3.5 cm, costae up to 12 cm apart, upper pinnae alternate, with stalk up to 5 mm; basal 1 or 2 pairs of pinnae larger, elliptic-lanceolate, up to 45 × 20 cm, lower part 2-pinnate, apex pinnatisect, acuminate; pinnules up to 15 pairs or more, spreading or nearly so, linear-lanceolate or broadly linear-lanceolate, up to 10 × 3 cm, apex long acuminate, pinnatisect, or pinnatipartite to pinnatisect, or pinnate, lower pinnules shortly stalked; secondary pinnules 1-5 pairs, lobes of secondary pinnules up to 10 pairs or more; secondary pinnules and lobes of secondary pinnules spreading, slightly oblique, or slightly falcate, ovate or elliptic-lanceolate, base broadly cuneate and adnate, pinnatilobate, dentate, or crenate, apex rounded or subtruncate; lobes of secondary pinnules slightly ascending, up to 6 pairs, mostly subfalcate, rectangular, shallowly repand or slightly shallowly crenate or serrate, truncate at apex; veins visible abaxially, not prominent adaxially, 1-3 pairs per lobe, veinlets simple. Lamina herbaceous, green abaxially, dark brown or gray-brown adaxially when dry; rachis, costae, and costules abaxially with brown, entire, lanceolate or linear, small scales, later glabrous. Sori orbicular, subbasal. Spores semicircular in equatorial view, perispore hyaline, with few rugate, vaguely granular projections.

Evergreen broad-leaved forests; 1400-2500 m. Yunnan (Gongshan, Jingdong, Yongde) [India, Nepal].

Cornopteris badia is the largest species with the most finely dissected lamina in the genus.

Plants having fronds with rachis, costae, and costules abaxially with sparse, brown, entire, lanceolate to linear, small scales and hyaline unicellular glandular hairs, and sharply serrate pinnule lobes have been named Cornopteris badia f. quadripinnatifida (M. Kato) W. M. Chu (Fl. Reipubl. Popularis Sin. 3(2): 365. 1999; C. quadripinnatifida M. Kato, Acta Phytotax. Geobot. 30(4-6): 114. 1979; C. badia var. pubescens Z. R. Wang; 毛复叶角蕨 mao fu ye jiao jue): evergreen broad-leaved forests; 1400-2500 m. Yunnan [India, Nepal].