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FOC | Family List | FOC Vol. 7 | Menispermaceae

19. Cyclea Arnott ex Wight, Ill. Ind. Bot. 1: 22. 1840.

轮环藤属 lun huan teng shu

Paracyclea Kudô & Yamamoto.

Vines. Petiole usually long; leaf blade peltate, palmately veined. Inflorescences axillary, terminal or on old stems, thyrsoid, usually narrow, rarely broad and sparse; bracts small. Male flowers: sepals usually 4 or 5(or 6), usually connate, 4- or 5-lobed, rarely free; petals 4 or 5, usually connate, entire or 4-8-lobed, rarely free, sometimes absent; stamens 4 or 5, connate into a peltate synandrium, anthers dehiscing transversely. Female flowers: sepals and petals 1 or 2, opposite to each other, rarely absent; carpel 1, stigma short, 3- or many lobed. Drupes obovate-globose to rotund, often slightly flat, style scar near base; endocarp bony, horseshoe-shaped, abaxially bearing 2 or 3 rows of tubercles on both surfaces of rib; condyle usually 1 or 2 cavities, often perforate between style scar and base. Seed horseshoe-shaped; embryo terete, embedded in endosperm.

About 29 species: S and SE Asia; 13 species (five endemic) in China.


1 Inflorescences cauliflorous or on leafless old branches, broad and large, many branched.   13 C. polypetala
+ Inflorescences axillary on normal leafy shoots, narrow and few branched (except C. barbata)   (2)
       
2 (1) Male inflorescences broad and large, many branched.   1 C. barbata
+ Male inflorescences narrow, thyrsoid, few branched, sometimes spikelike   (3)
       
3 (2) Male flowers with sepals conspicuously connate   (4)
+ Male flowers with sepals free or only connate at base   (7)
       
4 (3) Male sepals forming cup, lobes ca. as long as tube; leaves and branchlets hispid; petioles not peltately inserted.   10 C. meeboldii
+ Male sepals not forming cup, lobes much shorter than tube; leaves and branchlets puberulent or subglabrous; petioles ± peltately inserted   (5)
       
5 (4) Male sepals urceolate, pubescent.   12 C. tonkinensis
+ Male sepals mitriform, glabrous   (6)
       
6 (5) Synandrium not exserted; male sepals 2.5-4 mm.   5 C. racemosa
+ Synandrium exserted; male sepals ca. 2 mm or shorter.   11 C. debiliflora
       
7 (3) Rachis and drupes glabrous   (8)
+ Rachis and drupes pubescent to hispid   (10)
       
8 (7) Endocarp ca. 7 mm.   2 C. sutchuenensis
+ Endocarp less than 5.5 mm   (9)
       
9 (8) Endocarp subtruncate or slightly emarginate; leaves 2.5-7 × 1.5-4 cm, glabrous or sparsely pilose, not papillate.   3 C. hypoglauca
+ Endocarp tapered at base; leaves 6-12 × 3-5.5 cm, abaxially densely setose or papillate.   4 C. wattii
       
10 (7) Leaf blade with margins coarsely 1- or 2-toothed.   9 C. ochiaiana
+ Leaf blade with margins entire or sparsely serrate   (11)
       
11 (10) Female flowers with petals as long as sepals; male flowers with petal 1 or absent.   8 C. gracillima
+ Female flowers with petals much smaller than sepals (female flowers unknown in C. longgangensis)   (12)
       
12 (11) Male sepals mitriform, 2.5-4 mm.   5 C. racemosa
+ Male sepals not mitriform   (13)
       
13 (12) Leaf margin long pubescent; male sepals oblong-elliptic.   6 C. longgangensis
+ Leaf margin shortly puberulent; male sepals obovate or spatulate.   7 C. insularis

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