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FOC | Family List | FOC Vol. 9 | Rosaceae

39. FRAGARIA Linnaeus, Sp. Pl. 1: 494. 1753.

草莓属 cao mei shu

Li Chaoluan (Li Chao-luang); Hiroshi Ikeda, Hideaki Ohba

Herbs perennial, mostly stoloniferous, polygamo-dioecious, usually spreading or appressed hairy. Stolons often rooting at nodes and forming plantlets. Leaves alternate, petiolate; stipules adnate to base of petiole, often membranous, sheathing; leaf blade 3-foliolate or pinnately 5-foliolate. Inflorescence erect, cymose or corymbiform, few flowered, rarely a solitary flower. Hypanthium obconic or turbinate. Sepals 5, valvate, persistent; epicalyx segments 5, alternating with and smaller than sepals, margin entire. Petals 5, white, rarely yellow, broadly obovate or suborbicular. Stamens numerous; anther 2-loculed. Carpels numerous, free, borne on convex receptacle; ovule ascending from middle of locule; style adaxial, short, persistent. Aggregate fruit formed from enlarged receptacle, berry-like, long conic to globose, fleshy. Achenes numerous, seated in pits on surface of aggregate fruit, minute, brittle. Seed testa membranous; cotyledons convex. x = 7.

About 20 species: temperate and subtropical regions of N hemisphere, extending to South America; nine species (three endemic, one introduced) in China.

1 Stems and petioles appressed hairy.   (2)
+ Stems and petioles spreading hairy.   (5)
2 (1) Sepals appressed to agregate fruit in fruit.   9 F. nubicola
+ Sepals reflexed in fruit.   (3)
3 (2) Leaflets 5; aggregate fruit ovoid.   3 F. pentaphylla
+ Leaflets 3, or if 5 then aggregate fruit globose or ellipsoid.   (4)
4 (3) Aggregate fruit long conic, ovoid, or rarely fusiform; epicalyx segments oblong, apex 2- or 3-lobed; leaflets 3.   7 F. daltoniana
+ Aggregate fruit globose or ellipsoid; epicalyx segments linear-lanceolate or linear, apex entire or lobed; leaflets 3 or 5.   8 F. gracilis
5 (1) Pedicels appressed hairy.   1 F. vesca
+ Pedicels densely spreading hairy.   (6)
6 (5) Sepals spreading or reflexed in fruit.   (7)
+ Sepals appressed to aggregate fruit in fruit.   (8)
7 (6) Leaflets 3, thin, both surfaces hairy, abaxially more densely so on veins; sepals spreading in fruit.   2 F. orientalis
+ Leaflets 5, thick, abaxially sparsely pilose, adaxially glabrous; sepals reflexed in fruit.   3 F. pentaphylla
8 (6) Leaflets (3 or)5, thin; plants silvery hairy throughout.   6 F. moupinensis
+ Leaflets 3, thick; plants brown-yellow hairy throughout.   (9)
9 (8) Aggregate fruit 1–1.5 cm in diam.; plants wild.   4 F. nilgerrensis
+ Aggregate fruit to 3 cm in diam.; plants cultivated.   5 F. ×ananassa

  • List of lower taxa


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