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FOC | Family List | FOC Vol. 17 | Lamiaceae

9. Gomphostemma Bentham, Edward's Bot. Reg. 15: t. 1292. 1830.

锥花属 zhui hua shu

Shrubs or perennial herbs. Stems with stellate hairs. Leaves petiolate; leaf blade margin serrate, adaxially stellate puberulent/hirsute, abaxially densely stellate lanate. Inflorescences axillary, sometimes basal, rarely terminal, cymes pedunculate or sessile, sometimes in spikes or panicles; bracts subcircular to linear-lanceolate. Calyx campanulate to tubular, 10-veined, 5-toothed, teeth equal or subequal. Corolla purple-red or yellow to white, 2-lipped, straight or curved, long exserted; tube abruptly or gradually dilated at throat, not hairy annulate inside; upper lip straight, slightly galeate, entire or emarginate, sometimes mucronulate; lower lip spreading, 3-lobed, middle lobe longest. Stamens 4, underlying upper lip, anterior 2 longest; filaments complanate, hairy on two sides or glabrous; anthers close together in pairs, cells parallel, transverse. Style included, apically subequally 2-cleft. Nutlets 1-4 maturing per flower, obovoid to ovoid, rarely globose, rugulose, glabrous or hirsute, drupelike, areole oblique.

About 36 species: China, India, Indonesia, Malaysia, Myanmar, Philippines, Thailand, Vietnam; 15 species in China.

Gomphostemma is one of few tropical rain forest genera in the Lamiaceae.


1 Inflorescences clearly pedunculate, inserted near base of stem.   (2)
+ Inflorescences often subsessile, inserted at mid and upper stem leaf axils.   (4)
       
2 (1) Bracts linear to capillary, 0.3-1.5 mm wide   6 Gomphostemma pseudocrinitum
+ Bracts lanceolate to ovate, 5-15 mm wide.   (3)
       
3 (2) Peduncle 2.5-7 cm; bracts purplish red, brownish purple when dry   15 Gomphostemma pedunculatum
+ Peduncle 0.2-2.2 cm; bracts greenish   14 Gomphostemma chinense
       
4 (1) Inflorescence an elongated thyrse of decussate, 1- or 2-flowered cymes.   (5)
+ Inflorescence a many-flowered cyme or group of cymes, often forming dense capitula.   (6)
       
5 (4) Calyx ca. 9 mm, teeth 3-3.2 mm; corolla 3.5-3.7 cm   1 Gomphostemma deltodon
+ Calyx 5-7 mm, teeth ca. 1 mm; corolla 1.6-2 cm   2 Gomphostemma microdon
       
6 (4) Bracts 0.5-3 mm wide.   (7)
+ Bracts 3-18 mm wide.   (9)
       
7 (6) Cymes 3- or 4-flowered; calyx teeth 8-9 mm   4 Gomphostemma leptodon
+ Cymes many flowered; calyx teeth 3-7 mm.   (8)
       
8 (7) Bracts 1- or 3-veined; calyx teeth linear to triangular-lanceolate   3 Gomphostemma parviflorum
+ Bracts inconspicuously 1- or 3-veined; calyx teeth linear   5 Gomphostemma crinitum
       
9 (6) Calyx teeth triangular, obtuse to short needlelike.   (10)
+ Calyx teeth triangular-lanceolate to linear-lanceolate, ± acuminate.   (11)
       
10 (9) Calyx teeth needlelike; nutlets compressed obovate, 3.5-4 mm wide, ± stellate at maturity   8 Gomphostemma lucidum
+ Calyx teeth obtuse; nutlets oblong triquetrous, 3-3.5 mm wide, glabrous at maturity   9 Gomphostemma arbusculum
       
11 (9) Leaf blade adaxially densely appressed tawny hirsute, abaxially with stipitate, tawny, stellate pubescence; corolla purple-red   11 Gomphostemma stellatohirsutum
+ Leaves sparsely pubescent or stellate hirsute/villous, abaxially stellate tomentose; corolla white to yellow (not recorded for G. callicarpoides).   (12)
       
12 (11) Leaves (7-)18-30 × (4-)7-14.5 cm, adaxially sparsely pubescent or minutely hispid.   (13)
+ Leaves 4-16 × 2-7 cm, adaxially sparsely stellate hirsute.   (14)
       
13 (12) Bracts 3-3.5 mm wide; calyx teeth subacuminate   10 Gomphostemma latifolium
+ Bracts 7-14 mm wide; calyx teeth caudate-acuminate   13 Gomphostemma sulcatum
       
14 (12) Leaf blade adaxially sparsely stellate hirsute   12 Gomphostemma callicarpoides
+ Leaf blade adaxially densely stellate pubescent   7 Gomphostemma hainanense

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