1 |
Leaves basal, appressed to substrate, often 2, rarely 1 or more than 2, often cordate, orbicular, or ovate |
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(2) |
+ |
Leaves cauline, well spaced or tufted, never appressed to substrate, various in shape but never cordate nor orbicular |
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(15) |
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2 (1) |
Petals simple |
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(3) |
+ |
Petals bilobed, anterior lobe sometimes small and obscure |
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(9) |
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3 (2) |
Flowers white; sepals ciliate or not |
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(4) |
+ |
Flowers green or greenish white; sepals not ciliate |
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(5) |
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4 (3) |
Leaves 3 or 4(-6), in a rosette; lip with a conic appendage at base; spur longer than pedicel and ovary. |
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1 H. delavayi |
+ |
Leaves 2, subopposite; lip lacking an appendage; spur shorter than pedicel and ovary. |
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2 H. siamensis |
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5 (3) |
Spur 8-13 mm, ca. as long as or longer than ovary; leaves 2-3 × as long as wide. |
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5 H. humidicola |
+ |
Spur indistinct or 1-7 mm, shorter than ovary; leaves slightly longer than wide |
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(6) |
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6 (5) |
Stem and ovary pubescent. |
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3 H. diplonema |
+ |
Stem and ovary glabrous |
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(7) |
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7 (6) |
Leaves 2, subopposite; spur ca. 7 mm. |
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6 H. diphylla |
+ |
Leaves 1 or 2, not as above; spur less than 2 mm |
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(8) |
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8 (7) |
Inflorescence 10-20-flowered; spur ca. 1.5 mm. |
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4 H. acianthoides |
+ |
Inflorescence 4- or 5-flowered; spur indistinct or absent. |
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7 H. reniformis |
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9 (2) |
Leaves with yellowish white markings adaxially |
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(10) |
+ |
Leaves lacking yellowish white markings adaxially |
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(11) |
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10 (9) |
Leaves thinly textured; sepals ciliate; petals deeply 2-lobed, lower lobe linear, 9(-13) mm. |
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8 H. fargesii |
+ |
Leaves relatively thickly textured; sepals not ciliate; petals shallowly 2-lobed, lower lobe ca. 0.5 mm. |
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9 H. wolongensis |
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11 (9) |
Dorsal sepal 3-6(-7) mm; petals not ciliate |
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(12) |
+ |
Dorsal sepal 7-13 mm; petals ciliate |
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(13) |
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12 (11) |
Spur 6-9 mm, shorter than pedicel and ovary. |
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10 H. aitchisonii |
+ |
Spur 11-13 mm, longer than pedicel and ovary. |
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11 H. balfouriana |
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13 (11) |
Leaves glaucous adaxially; base of peduncle densely pubescent; dorsal sepal 10-13 mm; spur nearly as long as pedicel and ovary. |
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12 H. glaucifolia |
+ |
Leaves green adaxially; base of peduncle sparsely pubescent or glabrous; dorsal sepal 7-9 mm; spur longer than pedicel and ovary |
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(14) |
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14 (13) |
Leaves with 5-7 white veins adaxially; lip lacking an appendage in mouth of spur. |
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13 H. tibetica |
+ |
Leaves with 5-7 green veins adaxially; lip with a conic appendage in mouth of spur. |
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14 H. szechuanica |
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15 (1) |
Petals bilobed |
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(16) |
+ |
Petals simple |
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(23) |
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16 (15) |
Petals shallowly bilobed; lip ± cross-shaped |
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(17) |
+ |
Petals deeply bilobed; lip not cross-shaped |
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(18) |
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17 (16) |
Flowers small; petals ca. 4 mm; lateral lobes of lip obovate-oblanceolate, spreading at a right angle to mid-lobe; spur 1.4-1.5 cm, ca. as long as pedicel and ovary. |
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15 H. schindleri |
+ |
Flowers rather large, petals 5-5.5 mm; lateral lobes of lip linear, spreading at an acute angle to mid-lobe; spur 2.5-3.5 cm, longer than pedicel and ovary. |
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16 H. linearifolia |
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18 (16) |
Petals usually with 6 or 7 filiform laciniae; lip with many filiform laciniae. |
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17 H. polytricha |
+ |
Petals and lip lacking filiform laciniae |
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(19) |
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19 (18) |
Sepals long acuminate or caudate-aristate at apex |
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(20) |
+ |
Sepals neither long acuminate nor caudate-aristate at apex |
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(21) |
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20 (19) |
Petals with anterior (lower) lobe ca. 2.5 mm, shorter than posterior (upper) lobe. |
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18 H. stenopetala |
+ |
Petals with anterior (lower) lobe 18-25 mm, longer than posterior (upper) lobe. |
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19 H. pantlingiana |
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21 (19) |
Petals and lip not ciliate on margins of lobes; petal lobes of unequal size, with posterior lobe shorter and wider than anterior lobe. |
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20 H. coultousii |
+ |
Petals and/or lip ciliate, or lip pubescent along apical margins of lobes; petal lobes of equal size |
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(22) |
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22 (21) |
Petals and lip with ciliate lobes; spur not lobed. |
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21 H. petelotii |
+ |
Petals lacking ciliate lobes; spur deeply bilobed at apex. |
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22 H. hosokawae |
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23 (15) |
Lateral lobes of lip with many long laciniae along outside margins |
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(24) |
+ |
Lateral lobes of lip sometimes serrate but lacking long laciniae along outside margins |
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(31) |
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24 (23) |
Petals linear, ca. 0.5 mm wide; spur thickened and geniculate-curved near middle. |
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23 H. medioflexa |
+ |
Petals not linear, 3-8 mm wide; spur not as above |
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(25) |
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25 (24) |
Petals falcate, subovate, ca. 8 mm wide |
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(26) |
+ |
Petals oblong or lanceolate, 4-5 mm wide |
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(28) |
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26 (25) |
Sepals not ciliate; connective ca. 8 mm wide; staminodes spatulate, ca. 2 mm; spur shorter than ovary. |
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28 H. yuana |
+ |
Sepals ciliate; connective 2-4 mm wide; staminodes elliptic, ca. 1 mm; spur ca. as long as or longer than ovary |
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(27) |
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27 (26) |
Inflorescence 1-4-flowered; spur 7-8.5 cm, much longer than ovary; connective ca. 2 mm wide. |
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29 H. intermedia |
+ |
Inflorescence 5- or more flowered; spur 3.5-4 cm, ca. as long as or slightly longer than ovary; connective 3-4 mm wide. |
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30 H. arietina |
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28 (25) |
Spur 4.5-6.5 cm, much longer than pedicel and ovary. |
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26 H. davidii |
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Spur 1.7-3.5 cm, shorter than or nearly as long as pedicel and ovary |
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(29) |
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29 (28) |
Connective 10-12 mm wide. |
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24 H. limprichtii |
+ |
Connective 2-5 mm wide |
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(30) |
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30 (29) |
Petals ± hairy adaxially; spur 17-18 mm; connective ca. 2 mm wide. |
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25 H. pectinata |
+ |
Petals glabrous adaxially; spur 20-35 mm; connective ca. 5 mm wide. |
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27 H. mairei |
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31 (23) |
Lateral sepals strongly oblique |
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(32) |
+ |
Lateral sepals not or slightly oblique |
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(40) |
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32 (31) |
Lip with a ring-shaped or cupular structure, or sometimes a lamella near entrance of spur |
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(33) |
+ |
Lip lacking a ring-shaped, cupular structure or lamella near entrance of spur |
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(36) |
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33 (32) |
Lateral lobes of lip much shorter than mid-lobe |
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(34) |
+ |
Lateral lobes of lip nearly as long as mid-lobe |
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(35) |
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34 (33) |
Spur 10-17 mm, shorter than pedicel and ovary; petals oblong, 3-4 mm. |
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36 H. acuifera |
+ |
Spur 20-29 mm, longer than pedicel and ovary; petals narrowly ovate or narrowly elliptic, 4-5 mm. |
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37 H. linguella |
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35 (33) |
Flowers white, with a brown vein on lateral sepals; spur 20-22 mm, slightly longer than pedicel and ovary. |
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38 H. rostellifera |
+ |
Flowers red-orange, with pale green sepals, without a brown vein on lateral sepals; spur 11-12 mm, shorter than pedicel and ovary. |
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39 H. rostrata |
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36 (32) |
Dorsal sepal much smaller than lateral sepals |
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(37) |
+ |
Dorsal sepal not or slightly smaller than lateral sepals |
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(38) |
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37 (36) |
Lateral sepals obliquely axe-shaped, ca. 7 × 13 mm; petals ca. 4 mm wide; spur 5-8 cm, pendulous, longer than pedicel and ovary; ovary glabrous. |
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31 H. commelinifolia |
+ |
Lateral sepals obliquely triangular, ca. 5 × 8 mm; petals ca. 1 mm wide; spur 2-2.2 cm, ca. as long as or shorter than pedicel and ovary; ovary pubescent. |
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32 H. austrosinensis |
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38 (36) |
Dorsal sepal with 3 serrulate keels on abaxial surface. |
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33 H. ciliolaris |
+ |
Dorsal sepal without keels on abaxial surface |
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(39) |
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39 (38) |
Rachis villous on ridges; petals triangular-lanceolate, ca. 6 mm; lip with a conic callus at base; spur 1.6-2 cm, ca. as long as pedicel and ovary. |
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34 H. hystrix |
+ |
Rachis glabrous; petals linear-lanceolate, 13-15 mm; lip without callus at base; spur 3-6 cm, much longer than pedicel and ovary. |
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35 H. fordii |
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40 (31) |
Flowers orange to red; lip mid-lobe bilobed. |
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40 H. rhodocheila |
+ |
Flowers not orange to red; lip or its mid-lobe simple |
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(41) |
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41 (40) |
Lip often unlobed; spur often absent. |
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41 H. malintana |
+ |
Lip deeply 3-lobed; spur present |
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(42) |
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42 (41) |
Lateral lobes of lip broad, subrhombic or suborbicular, apical margin serrate |
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(43) |
+ |
Lateral lobes of lip narrow, not as above, margins never serrate |
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(44) |
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43 (42) |
Plants 9-25 cm tall; leaves cordate or ovate; spur 17-20 mm, slightly shorter or nearly as long as pedicel and ovary. |
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42 H. finetiana |
+ |
Plants 35-87 cm tall; leaves oblong to narrowly elliptic; spur to 4 cm, longer than pedicel and ovary. |
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43 H. dentata |
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44 (42) |
Mid-lobe of lip suberect or curved upward, apex adnate to tip of hood formed by dorsal sepal and petals |
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(45) |
+ |
Mid-lobe of lip never suberect or curved upward, apex never adnate to hood formed by dorsal sepal and petals |
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(46) |
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45 (44) |
Floral bracts shorter than pedicel and ovary; lateral sepals reflexed; spur cylindric, to 21 mm, much longer than pedicel and ovary. |
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44 H. lucida |
+ |
Floral bracts longer than flower; lateral sepals spreading, not reflexed; spur clavate, 3-6 mm, shorter than pedicel and ovary. |
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45 H. shweliensis |
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46 (44) |
Leaves tufted at base of stem; spur longer than ovary |
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(47) |
+ |
Leaves not tufted at base of stem; spur longer or shorter than ovary |
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(50) |
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47 (46) |
Leaves narrowly elliptic or broadly oblanceolate, 18-20 mm wide; flowers pale green. |
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46 H. iyoensis |
+ |
Leaves linear or linear-lanceolate, 4-15 mm wide; flowers yellowish green or white |
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(48) |
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48 (47) |
Leaves 4 or 5, 4-6 mm wide; dorsal sepal ca. 2 mm; spur narrowly cylindric. |
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47 H. viridiflora |
+ |
Leaves 6-13, 8-15 mm wide; dorsal sepal ca. 5 mm; spur narrowly cylindric-clavate |
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(49) |
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49 (48) |
Dorsal sepal broadly ovate; lateral sepals narrowly ovate; petals ovate-ligulate; lip with lateral lobes narrower than mid-lobe. |
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48 H. plurifoliata |
+ |
Dorsal sepal elliptic; lateral sepals elliptic; petals linear; lip with lateral lobes and mid-lobe equal in width. |
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49 H. tonkinensis |
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50 (46) |
Dorsal sepal 7-8 mm |
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(51) |
+ |
Dorsal sepal 3-4 mm |
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(52) |
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51 (50) |
Spur narrowly cylindric, 3-5 mm, shorter than pedicel and ovary; dorsal sepal narrowly ovate, ca. 4 mm wide; petals ca. 2.5 mm wide; flowers yellow. |
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50 H. fulva |
+ |
Spur clavate, 8-13 mm, ca. as long as pedicel and ovary; dorsal sepal cordate or broadly ovate, 5-6 mm wide; petals obliquely ovate-triangular, 3-3.8 mm wide; flowers yellowish green. |
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51 H. marginata |
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52 (50) |
Floral bracts longer than ovary; flowers pale yellowish green. |
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52 H. leptoloba |
+ |
Floral bracts shorter or nearly as long as ovary; flowers pinkish white or pale green |
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(53) |
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53 (52) |
Floral bracts much shorter than pedicel and ovary; flowers pinkish white; lateral sepals 2-veined; petals obliquely oblong-ovate; spur ca. 21 mm. |
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53 H. nematocerata |
+ |
Floral bracts nearly as long as pedicel and ovary; flowers pale green; lateral sepals 3-veined; petals lanceolate; spur ca. 17 mm. |
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54 H. furcifera |