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4. Holboellia Wallich, Tent. Fl. Napal. 1: 23. 1824.
八月瓜属 ba yue gua shu
Woody climbers, evergreen or deciduous. Monoecious. Stems twining. Winter bud outer scales numerous, seriate. Leaves alternate, usually long petiolate; leaf blade palmately 3--9-foliolate or pinnately 3-foliolate; petiolules usually unequal with terminal one longest; leaflet margins entire. Inflorescences axillary, clustered, corymbiform racemes, rarely elongated racemes; several to many flowered. Sepals 6, greenish white or purple, oblong, slightly fleshy, outer 3 valvate, inner 3 usually smaller. Petals 6, minute, opposite stamens, rudimentarily nectariferous. Male flowers: stamens 6, free; connective apically appendaged or unappendaged. Pistillodes 3, minute. Female flowers: carpels 3, straight. Stigma terminal or oblique, often cleft. Fruit oblong to ellipsoid, fleshy, indehiscent. Seeds embedded in pulp.
Twenty species: SE Asia, China, Himalayas; nine species (five endemic) in China.
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| 1 |
Leaves pinnately 3-foliolate. |
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(2) |
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Leaves palmately 3--7(--9)-foliolate. |
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(4) |
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| 2 (1) |
Leaflets papery; outer sepals of male flower 1.6--1.8 cm |
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3 H. brachyandra |
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Leaflets subleathery to thick leathery; outer sepals of male flower to 1 cm. |
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(3) |
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| 3 (2) |
Leaflets thick leathery, abaxially farinaceous green; outer sepals of male flower to 1 cm |
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1 H. coriacea |
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Leaflets leathery to subleathery, abaxially pale green; outer sepals of male flower 3--7 mm |
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2 H. parviflora |
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| 4 (1) |
Stems and branches angular or narrowly winged. |
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(5) |
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Stems and branches terete, striate, neither angular nor winged. |
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(6) |
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| 5 (4) |
Veins of leaflets conspicuously impressed on adaxial surface; outer sepals of male flower 0.7--0.8 cm |
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8 H. chapaensis |
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Veins of leaflets prominent on both surfaces, sometimes plane but never impressed on adaxial surface; outer sepals of male flower 1.2--1.3 cm |
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9 H. pterocaulis |
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| 6 (4) |
Leaflets 4--14 × as long as wide (less than 2 × in subsp. obtusa) |
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5 H. angustifolia |
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Leaflets 1.5--2.5 × as long as wide. |
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(7) |
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| 7 (6) |
Outer sepals of male flower ca. 0.8 × 0.3 cm; petals orbicular |
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7 H. medogensis |
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Outer sepals of male flower 1.1--2.2 × 0.4--1 cm; petals not orbicular (sometimes suborbicular in H. grandiflora). |
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(8) |
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| 8 (7) |
Stamens ca. 1.5 cm, filaments ca. 1 cm; seeds black |
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4 H. grandiflora |
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Stamens 0.8--1.2 cm, filaments 0.5--7 cm; seeds brown |
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6 H. latifolia |
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List of lower taxa
| Related Links (opens in a new window) |
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Treatments in Other Floras @ www.efloras.org
Other Databases
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