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FOC | Family List | FOC Vol. 19 | Annonaceae

5. Miliusa Leschenault ex A. Candolle, Mém. Soc. Phys. Genève. 5: 213. 1832.

野独活属 ye du huo shu

Authors: Bingtao Li & Michael G. Gilbert

Saccopetalum Bennett.

Trees or erect shrubs. Inflorescences axillary or superaxillary, fasciculate or cymose, 1-flowered. Flowers mostly bisexual. Pedicel slender, usually elongate. Torus elevated, usually long pubescent. Sepals 3, small, valvate, connate at base. Petals 6, in 2 whorls, with each whorl valvate; outer petals similar to sepals; inner petals large, erect, thin, base shallowly saccate and shortly clawed, margin initially cohering but readily free, apex usually revolute. Stamens many; connectives narrow and not concealing anther locules, apex subapiculate. Carpels many, oblong; ovules 1-8 per carpel; stigmas capitate, ovoid, or globose. Fruit apocarpous; monocarps many, subsessile to long stipitate, globose to ovoid. Seeds 1 to several per monocarp.

About 38 species: tropical and subtropical Asia and Australia; seven species (three endemic) in China.

1 Leaf blade broadest above middle; ovules 1 per carpel   (2)
+ Leaf blade broadest at or below middle; ovules 2-8 per carpel   (3)
2 (1) Leaf blade apex acuminate to shortly acuminate, secondary veins obscure and 7-9 on each side of midvein; pedicel 1-1.4 cm.   1 M. tenuistipitata
+ Leaf blade apex obtuse, secondary veins raised and 12-15 on each side of midvein; pedicel 2-3 cm.   2 M. cuneata
3 (1) Shrubs; leaf blade glabrous or with larger veins only sparsely puberulent and glabrescent; sepals and petals sparsely pubescent   (4)
+ Trees; leaf blade abaxially pubescent or tomentose; sepals and petals pubescent or tomentose   (5)
4 (3) Leaf blade secondary veins 13-20 on each side of midvein; pedicel 0.5-0.7 cm, superaxillary; inner petals ca. 0.3 cm.   3 M. glochidioides
+ Leaf blade secondary veins 10-12 on each side of midvein; pedicel 4-6.5 cm, axillary; inner petals ca. 1.8 cm.   4 M. balansae
5 (3) Leaf blade 5-10 cm wide, base obliquely cordate; flowers 3-6 in leaf-opposed cymes or on short and few-leaved branches; carpels glabrous.   5 M. velutina
+ Leaf blade 1.8-5 cm wide, base obtuse to rounded and slightly oblique; flowers 1-3 and axillary; carpels pubescent   (6)
6 (5) Pedicel 3.5-7.5 cm; outer petals as long as sepals; ovules 2 per carpel; monocarps glabrous.   6 M. sinensis
+ Pedicel 1.5-3 cm; outer petals longer than sepals; ovules 8 per carpel; monocarps puberulent.   7 M. horsfieldii

Lower Taxa


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