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FOC | Family List | FOC Vol. 8 | Brassicaceae

94. Neotorularia Hedge & J. Léonard, Bull. Jard. Bot. Belg. 56: 393. 1986.

念珠芥属 nian zhu jie shu

Torularia (Cosson) O. E. Schulz in Engler, Pflanzenr. 86(IV. 105): 213. 1924, not Bonnemaison (1828); based on Sisymbrium Linnaeus sect. Torularia Cosson, Comp. Fl. Atlant. 2: 136. 1887.

Herbs annual, biennial, or perennial with a woody caudex. Trichomes stalked or subsessile, 1- or 2-forked sometimes also simple. Stems erect or prostrate, usually several from base. Basal leaves petiolate or sessile, rosulate or not, entire, dentate, or pinnatisect. Cauline leaves often similar to basal ones, rarely absent. Racemes few to several flowered, ebracteate or rarely bracteate. Fruiting pedicels erect, ascending, or divaricate. Sepals ovate or oblong, erect or spreading, rarely reflexed, glabrous or pubescent, base of lateral pair not saccate. Petals white or pink, sometimes yellowish, longer or rarely shorter than sepals; blade obovate, spatulate, or oblanceolate, apex obtuse or retuse; claw distinct or not. Stamens 6, tetradynamous; filaments rarely dilated at base; anthers ovate or narrowly oblong, apex apiculate or obtuse. Median glands absent; lateral ones semilunar or 1 on each side of lateral stamen. Ovules (8-)16-36(-44) per ovary. Fruit dehiscent siliques, linear or rarely linear-oblong, terete or 4-angled, sessile; valves with branched trichomes sometimes mixed with simple ones, rarely glabrescent, torulose; replum rounded; septum complete; style obsolete or to 1 mm; stigma capitate, entire or 2-lobed. Seeds uniseriate, wingless, oblong or ovate, plump; seed coat reticulate, not mucilaginous when wetted; cotyledons incumbent.

About 14 species: C and SW Asia, one species extending into North America and another into N Africa and Europe; six species (one endemic) in China.

1 Perennials, at least lowermost portion of raceme bracteate.   (2)
+ Annuals or biennials, if perennial then style more than 1 mm; racemes ebracteate.   (3)
2 (1) Fruit (9-)12-25(-32) × 0.5-0.9(-1) mm, uniform in width; petals 3-5(-8) mm; seeds uniseriate; racemes bracteate along lowermost portion, rarely throughout   1 N. humilis
+ Fruit 3-10(-15) × 1-1.3 mm, widest near or at base; petals 1-2.5(-3) mm; seeds often subbiseriate at basal portion of fruit; racemes bracteate throughout or rarely only along proximal half   2 N. brachycarpa
3 (1) Racemes rachis flexuous; fruit straight; Nei Mongol   6 N. qingshuiheense
+ Racemes rachis straight; fruit twisted or curved apically; elsewhere.   (4)
4 (3) Stem leafless; petals less than 2 mm; basal leaves including petiole less than 1 cm   5 N. brevipes
+ Stem leafy; petals more than 2 mm; basal leaves including petiole more than 1 cm.   (5)
5 (4) Fruiting pedicels narrower than fruit, (3-)4-6(-8) mm; petals 5-7 mm   3 N. korolkowii
+ Fruiting pedicels as thick as fruit, 0.3-1(-2) mm; petals (2-)2.5-3.5(-4) mm   4 N. torulosa

Lower Taxa


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