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FOC | Family List | FOC Vol. 12 | Rhamnaceae

1. Paliurus Miller, Gard. Dict. Abr., ed. 4. 1754.

马甲子属 ma jia zi shu

Trees or shrubs, small to medium-sized, evergreen or deciduous. Leaves alternate, 3-veined from base, margin subentire to serrate; stipules usually changed into 1 or 2 lignified, erect or revurved spines. Flowers bisexual, 5-merous, perigynous, few to many in axillary, pedunculate cymes. Pedicel short, often elongated in fruit. Calyx tube hemispheric to dish-shaped; sepals free, deltoid, 1.5-2.5 × 1.5-2.5 mm, abaxially ± pubescent to glabrous, adaxially keeled, apex acute to weakly acuminate. Petals spatulate or unguiculate,1-2 mm, often enfolding stamens. Stamens free, 1.5-2 mm; filaments subulate, thin; anthers ovoid, latrorse. Disk adnate with calyx tube, pentagonal or rounded, thick, fleshy, glabrous. Ovary semi-inferior, 2- or 3-loculed, with 1 ovule per locule, slightly immersed in disk; style cylindric or flattened, with 2 or 3 ± diverging stylar branches. Fruit a dry, indehiscent, disk- to cup-shaped or hemispheric, winged drupe, base ± distinctly conical, often with remains of calyx tube, apex rounded to ± flattened; mesocarp leathery to corky; endocarp lignified. Seeds brownish, shiny, obovoid to ovoid; endosperm scanty, cotyledons flat.

Five species: E Asia, Europe; five species (three endemic, one introduced) in China.

1 Drupes with a narrow, apically inserted wing; stipular spines 1 or 2 per node   (2)
+ Drupes with a broad, papery, medially inserted wing; stipular spines 2 per node   (3)
2 (1) Fruit glabrous, wing entire; branches with 1 recurved spine per node.   2 P. hirsutus
+ Fruit densely pubescent, wing ± distinctly 3-partite; branches with 2 erect spines per node.   1 P. ramosissimus
3 (1) Branches with 1 erect and 1 recurved spine per node.   5 P. spina-christi
+ Branches with exclusively erect, never hooklike recurved spines   (4)
4 (3) Fruit 1.4-2(-2.6) cm in diam.; petiole 3-5(-8) mm, pubescent.   4 P. orientalis
+ Fruit 2-3.5 cm in diam.; petiole (6-)8-20 mm, glabrous or subglabrous.   3 P. hemsleyanus

Lower Taxa


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