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FOC | Family List | FOC Vol. 17 | Lamiaceae

44. Paraphlomis (Prain) Prain, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74: 721. 1908.

假糙苏属 jia cao su shu

Phlomis Linnaeus sect. Paraphlomis Prain, Ann. Roy. Bot. Gard. (Calcutta) 9: 60. 1901.

Herbs or subshrubs, rhizomatous. Stems ascending or erect. Leaves sessile or long petiolate; leaf blade membranous to ± leathery, margin dentate. Verticillasters axillary; bracteoles lanceolate to bristlelike, sometimes minute, early deciduous. Calyx tubular to obconical, mouth sometimes slightly constricted, gradually or abruptly dilated, truncate or pleated; veins 5-10, ± distinct; teeth 5, broadly triangular to subulate, straight, spreading or reflexed, bristlelike-acuminate to acute or widened into wings from veins. Corolla 2-lipped; tube hairy annulate inside, gradually dilated at apex; upper lip flat, erect or galeate, densely hairy outside; lower lip spreading at about 90°, 3-lobed, middle lobe larger. Stamens 4, anterior 2 longer; filaments filiform, complanate, ± hairy; anther cells 2, parallel or divergent. Ovary apex truncate, glabrous or hairy. Style apex subequally 2-cleft, lobes subulate. Disc ringlike or cupulate, apex truncate. Nutlets obovoid or triquetrous oblong, glabrous or hairy.

About 24 species: China, India, Indonesia, Laos, Malaysia, Myanmar, Thailand, Vietnam; 23 species in China.

1 Calyx tubular or tubular-campanulate.   (2)
+ Calyx obconical.   (11)
2 (1) Calyx teeth minute, mouth subtruncate, pleated   10 Paraphlomis membranacea
+ Calyx teeth conspicuous, broadly triangular to subulate, apex bristlelike to acute.   (3)
3 (2) Leaves ovate, ovate-orbicular, or subcircular.   (4)
+ Leaves elliptic, elliptic-ovate, to oblong-elliptic.   (6)
4 (3) Stems and leaves sparsely bristly; calyx tube densely bristly, teeth narrowly triangular, apex bristlelike   7 Paraphlomis seticalyx
+ Stems, leaves, and calyces densely pubescent and/or villous; calyx teeth broadly triangular, apex acute.   (5)
5 (4) Erect herbs; stems, leaves, and calyces densely villous and pubescent; corolla densely white villous outside; bracteoles minute   8 Paraphlomis albiflora
+ Ascending herbs; stems, leaves, and calyces densely villous only; corolla sparsely pubescent outside; bracteoles linear, 2-5 mm   9 Paraphlomis foliata
6 (3) Calyx bristly.   (7)
+ Calyx not bristly.   (9)
7 (6) Leaves ± leathery; leaves and stem glabrous; calyx sparsely white bristly at apex and along veins otherwise strigose   4 Paraphlomis paucisetosa
+ Leaves membranous or thin papery; leaves and stem hairy; calyx densely bristly.   (8)
8 (7) Leaves thin papery; leaves and stem densely spreading bristly   5 Paraphlomis patentisetulosa
+ Leaves membranous, strigose and finely papillate except for hispid veins of adaxial surface; abaxially white granular, finely strigose and sparsely hispid; stem densely retrorse fine strigose   3 Paraphlomis hispida
9 (6) Stems and calyx densely tawny hirsute   2 Paraphlomis hirsutissima
+ Stems and calyx not tawny hirsute.   (10)
10 (9) Stems retrorse strigose; calyx glabrous or hirtellous becoming glabrescent in fruit   1 Paraphlomis javanica
+ Stems and calyx densely and finely bristly puberulent   6 Paraphlomis setulosa
11 (1) Calyx teeth lanceolate-triangular, acute.   (12)
+ Calyx teeth not lanceolate-triangular.   (15)
12 (11) Verticillasters surrounded by leaflike bracts, with linear basal bracteoles   11 Paraphlomis albotomentosa
+ Verticillasters surrounded by linear bracts, with bristlelike basal bracteoles.   (13)
13 (12) Leaves elliptic or elliptic-ovate; calyx densely tomentose outside   12 Paraphlomis tomentosocapitata
+ Leaves broadly ovate or ovate-lanceolate to lanceolate; calyx pubescent outside.   (14)
14 (13) Leaves 3-5 cm wide; calyx ca. 9.5 mm; corolla ca. 1.8 cm   13 Paraphlomis lancidentata
+ Leaves 6.5-9.5 cm wide; calyx ca. 11 mm; corolla ca. 1.4 cm   14 Paraphlomis reflexa
15 (11) Calyx teeth triangular or inconspicuous, calyx mouth subtruncate, distinctly extended into winged extensions from veins.   (16)
+ Calyx teeth broadly triangular to broadly ovate-triangular or subulate, apex acuminate to acute.   (17)
16 (15) Leaves oblong to ovate-oblong, 5-8 × 2.4-3.4 cm, margin undulate-crenate; calyx teeth conspicuously triangular, calyx mouth not subtruncate   22 Paraphlomis brevifolia
+ Leaves lanceolate to narrowly lanceolate, 7-15 × 1.5-3 cm, margin remotely, shallowly serrate above base; calyx teeth inconspicuous, calyx mouth subtruncate   23 Paraphlomis subcoriacea
17 (15) Apical part of stem, petiole, and abaxial leaf surface densely retrorse pilose, especially on leaf veins   19 Paraphlomis albida
+ Apical part of stem, petiole, and abaxial leaf surface not densely retrorse pilose.   (18)
18 (17) Plants sparsely pubescent or subglabrous   16 Paraphlomis lanceolata
+ Plants densely retrorse strigose or pubescent.   (19)
19 (18) Corolla tube shorter than calyx tube; calyx teeth subulate   15 Paraphlomis gracilis
+ Corolla tube longer than calyx tube; calyx teeth not subulate.   (20)
20 (19) Leaves lanceolate to sublanceolate.   (21)
+ Leaves ovate.   (22)
21 (20) Stems densely spreading pubescent toward base, conspicuously grooved; leaves lanceolate, apex long acuminate, base attenuate; petiole ca. 2 cm; corolla conspicuously hairy annulate inside; ovary glabrous   17 Paraphlomis parviflora
+ Stems densely pubescent throughout, not grooved; leaves oblong-elliptic, apex acute, base cuneate; petioles ca. 1 cm; corolla inconspicuously hairy annulate inside; apex of ovary puberulent   18 Paraphlomis kwangtungensis
22 (20) Leaves membranous; corolla tube much exserted; anthers ciliate; ovary glabrous   20 Paraphlomis intermedia
+ Leaves papery; corolla tube slightly exserted; anthers not ciliate; ovary apex pubescent   21 Paraphlomis pagantha

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