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37. SIBBALDIA Linnaeus, Sp. Pl. 1: 284. 1753.

山莓草属 shan mei cao shu

Li Chaoluan (Li Chao-luang); Hiroshi Ikeda, Hideaki Ohba

Sibbaldianthe Juzepczuk.

Herbs perennial, often woody based. Flowering stems prostrate to erect. Leaves pinnate or palmately 3–5-foliolate; leaflets serrate at apex or margin, rarely entire. Inflorescence a cyme or solitary flower. Flowers usually bisexual, rarely unisexual. Hypanthium saucer-shaped or cupular. Sepals (4 or)5, alternate, persistent; epicalyx segments (4 or)5. Petals (4 or)5, yellow, purple-red, or white. Disk usually markedly broad, rarely not so. Stamens (4 or)5(–10); anthers 2-loculed. Carpels 4–20, free; ovule usually ascending; style sub-basal, lateral, or subterminal. Achenes few, inserted on elevated, dry receptacle. x = 7.

About 20 species: arctic and alpine regions of N hemisphere; 13 species (four endemic) in China.


1 Radical leaves pinnately 3–11-foliolate.   (2)
+ Radical leaves palmately 3–5-foliolate.   (5)
       
2 (1) Leaflets abaxially sericeous, strigose, or subglabrous.   (3)
+ Leaflets abaxially densely white tomentose.   (4)
       
3 (2) Flowers 5-merous; petals yellow or white, longer than sepals; leaflets abaxially strigose, adaxially subglabrous, terminal one (2 or)3-serrate at apex.   10 S. adpressa
+ Flowers 4- or 5-merous; petals white, equaling or only slightly longer than sepals; leaflets appressed sericeous on both surfaces, margin entire.   11 S. sericea
       
4 (2) Flowering stems and petioles densely white tomentose; leaflets abaxially inconspicouously veined and densely white tomentose.   12 S. micropetala
+ Flowering stems and petioles yellow pilose and white tomentose; leaflets abaxially prominently veined.   13 S. phanerophlebia
       
5 (1) Radical leaves 5-foliolate.   (6)
+ Radical leaves 3-foliolate.   (8)
       
6 (5) Leaflets densely white sericeous on both surfaces.   9 S. omeiensis
+ Leaflets appressed pilose on both surfaces.   (7)
       
7 (6) Leaflets obovate-oblong or oblong, central and lateral ones subequal, apex 2–6-serrate; petals 5, purple-red, longer than sepals.   7 S. purpurea
+ Leaflets obovate, 2 lateral ones much smaller than 3 central ones, apex (2 or)3-serrate; petals 4(or 5), yellow-white, nearly equaling sepals.   8 S. pentaphylla
       
8 (5) Leaflet margin entire except for (2 or)3–5-dentate apex.   (9)
+ Leaflet margin serrate or lobed.   (11)
       
9 (8) Petals 5, shorter than sepals.   1 S. procumbens
+ Petals 4 or 5, nearly equaling or slightly longer than sepals.   (10)
       
10 (9) Leaflets broadly obovate, base rounded to broadly cuneate; flowers numerous, in cymes; petals 5, equaling sepals.   2 S. cuneata
+ Leaflets obovate-oblong, base cuneate; flowers 1(–3); petals 4, nearly equaling or slightly longer than sepals.   3 S. tetrandra
       
11 (8) Flower solitary, terminal; petals white, longer than sepals.   4 S. perpusilloides
+ Flowers 2–6, in cymes; petals red or purple-red, shorter than or nearly equaling sepals.   (12)
       
12 (11) Leaflets broadly obovate to suborbicular, 0.3–1.5 cm, both surfaces appressed pilose; petals red, nearly equaling sepals.   5 S. tenuis
+ Leaflets broadly elliptic, 1.3–2.5 cm, abaxially densely yellow hirsute; petals purple-red, slightly shorter than or equaling sepals.   6 S. sikkimensis

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