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5. Stauntonia de Candolle, Syst. Nat. 1: 511, 513. 1817.
野木瓜属 ye mu gua shu
Parvatia Decaisne.
Woody climbers, evergreen. Monoecious or dioecious. Winter bud scales many, in several whorls; outer scales imbricate, usually short and broad; inner scales longer than outer ones, ligulate to straplike. Leaves alternate, long petiolate; leaf blade palmately 3--9-foliolate or pinnately 3-foliolate; petiolules unequal; leaflet margins entire. Inflorescences axillary, several to more than 10-flowered; bracts subtending inflorescences, scaly. Sepals 6, outer 3 valvate. Petals absent or 6 and inconspicuous. Male flowers: stamens 6; filaments connate into a tube, basally, or only at base; connective apical appendage often hornlike or apiculate. Pistillodes 3, concealed within filaments tube. Female flowers: sepals usually slightly larger than those of male flowers. Staminodes minute, squamose, sessile, inserted at base of carpels opposite petals. Carpels 3, straight, sessile; ovules in several rows. Stigma terminal. Fruit single or in pairs, rarely 3-fascicled, sometimes dehiscent along abaxial suture. Seeds numerous, embedded in pulp.
About 25 species: China, N India, Japan, Myanmar, Vietnam; 20 species (16 endemic) in China.
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1 |
Petals 6. |
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(2) |
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Petals absent. |
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(6) |
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2 (1) |
Leaves palmately 5--7-foliolate, sometimes 3- or 4-foliolate apically on branches. |
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(3) |
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Leaves pinnately 3-foliolate. |
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(4) |
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3 (2) |
Leaflets adaxially dull; petals 2--3 mm; fruit 4--6 × ca. 2.5 cm |
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4 S. maculata |
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Leaflets adaxially shiny; petals ca. 1.5 mm; fruit 7--10 × 3--5 cm |
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5 S. chinensis |
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4 (2) |
Branches, petiole, and petiolules narrowly winged; leaflets abaxially pulverulently or papillately puberulous; inflorescence 1- or 2-flowered; male flower outer 3 sepals 2.2--2.5 cm |
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1 S. decora |
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Branches, petiole, and petiolules not winged; leaflets glabrous; inflorescence many flowered; male flower outer 3 sepals 0.5--0.8 cm. |
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(5) |
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5 (4) |
Leaflets subleathery; secondary veins and tertiary veinlets conspicuously prominent on both surfaces; stamen connective apical appendage longer than anther |
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2 S. brunoniana |
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Leaflets papery; secondary veins and tertiary veinlets abaxially prominent, adaxially inconspicuous; stamen connective apical appendage as long as anther |
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3 S. elliptica |
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6 (1) |
Stamen connective unappendaged |
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20 S. leucantha |
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Stamen connective appendaged. |
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(7) |
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7 (6) |
Connective apical appendage 3--5 mm (sometimes ca. 2 mm in S. purpurea). |
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(8) |
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Connective apical appendage to 2.5 mm. |
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(12) |
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8 (7) |
Leaflets abaxially pale green |
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10 S. libera |
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Leaflets abaxially grayish green or glaucous. |
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(9) |
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9 (8) |
Leaflet base cuneate to broadly cuneate |
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9 S. duclouxii |
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Leaflet bases obtusely rounded, rounded, or slightly cordate. |
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(10) |
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10 (9) |
Leaflets abaxially whitish; pedicel ca. 1 cm; connective apical appendage ca. 4 mm |
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8 S. trinervia |
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Leaflets abaxially grayish green; pedicel 2--5 cm; connective apical appendage 2--3 mm. |
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(11) |
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11 (10) |
Leaflets 6--9.5 cm wide, abaxially not maculate, margin revolute when dry; sepals of male flowers ca. 1.8 cm |
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6 S. glauca |
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Leaflets 2.5--5 cm wide, abaxially densely whitish brown maculate, margin crisp when dry; sepals of male flowers 1--1.5 cm |
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7 S. purpurea |
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12 (7) |
Connective apical appendage 2--2.5 mm. |
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(13) |
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Connective apical appendage minute to 1 mm. |
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(15) |
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13 (12) |
Leaflets 12--17 cm, subleathery, apex with a ca. 3.5 cm cauda; outer 3 sepals of male flowers 2--2.5 cm |
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13 S. yaoshanensis |
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Leaflets less than 7 cm, leathery, apex not caudate; outer 3 sepals of male flowers 0.8--1.4 cm. |
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(14) |
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14 (13) |
Leaflet apex mucronate and apiculate; outer 3 sepals of male flowers ca. 8 cm |
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11 S. oligophylla |
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Leaflet apex emarginate to obcordate, not mucronate and apiculate; outer 3 sepals of male flowers ca. 1.4 cm |
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12 S. obcordatilimba |
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15 (12) |
Connective apical appendage ca. 1 mm. |
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(16) |
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Connective apical appendage scarcely visible to 0.6 mm. |
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(18) |
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16 (15) |
Leaflets 3(--5), abaxially whitish green, margin conspicuously revolute |
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14 S. conspicua |
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Leaflets 5--9, abaxially pale green or glaucous, margin not revolute. |
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(17) |
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17 (16) |
Leaflets obovate, obovate-oblong, or spatulate, abaxially not maculate |
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16 S. obovatifoliola |
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Leaflets oblanceolate, oblong-lanceolate, or oblong abaxially densely maculate |
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15 S. pseudomaculata |
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18 (15) |
Leaflets lanceolate-linear to lanceolate, abaxially densely maculate; connective apical appendage ca. 0.6 mm; plants dioecious |
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17 S. cavalerieana |
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Leaflets obovate, oblong, broadly elliptic, oblanceolate, or spatulate, abaxially not maculate; connective apical appendage scarcely visible; plants monoecious. |
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(19) |
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19 (18) |
Leaflets 3--5, obovate, oblong, broadly elliptic, or oblanceolate, apex rounded or sometimes acute to acuminate; sepals thin, not fleshy |
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18 S. obovata |
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Leaflets 5--9, spatulate, apex long caudate and with a filiform easily broken tip; sepals thick, subfleshy |
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19 S. brachyanthera |
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List of lower taxa
Related Links (opens in a new window) |
Treatments in Other Floras @ www.efloras.org
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