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28. Suaeda Forsskål ex J. F. Gmelin, Onomat. Bot. Compl. 8: 797. 1776.
[nom. cons.]
碱蓬属 jian peng shu
Herbs annual, subshrubs, or shrubs, sometimes glaucous, glabrous, rarely pubescent. Stem erect, obliquely spreading, or prostrate. Leaves usually sessile and linear, terete or semiterete, rarely clavate or slightly compressed, fleshy, margin entire. Flowers small, bisexual, sometimes also female, usually 3 to numerous in glomerules, these axillary or on dwarf, axillary branchlets, sometimes dwarf branchlet base fused to leaf base and glomerule thus appearing to arise from petiole; bractlets 2 per flower, white, scale-like, membranous. Perianth subglobose, hemispheric, or urceolate, 5-parted or -lobed, slightly fleshy or herbaceous; segments abaxially thickened or becoming winglike or hornlike, rarely not modified, adaxially concave or cucullate. Stamens 5; filaments short, flattened; anthers cylindric, ellipsoid, or subglobose, without an appendage. Ovary ovoid or globose; stigmas 2 or 3(-5), usually recurved, papillate throughout. Utricle enclosed by perianth; pericarp membranous, free from seed. Seed horizontal or vertical, lenticular, reniform, ovoid, or globose; testa thinly leathery or membranous; embryo green or whitish, planospiral, slender; perisperm scant or absent.
About 100 species: Asia, Europe, North America, and seashores worldwide: 20 species (two endemic) in China.
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1 |
Glomerule borne on dwarf, axillary branchlet, dwarf branchlet base fused with petiole and thus glomerule appearing to be borne on petiole. |
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(2) |
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Glomerule borne in leaf axil or on dwarf, axillary branchlet, dwarf branchlet base not fused with petiole. |
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(7) |
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2 (1) |
Subshrubs. |
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(3) |
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Herbs annual. |
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(4) |
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3 (2) |
Leaves usually 3-5 mm, abruptly contracted at base; perianth parted to middle. |
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1 S. microphylla |
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Leaves usually 5-15 mm, attenuate into short petiole at base; perianth parted to base. |
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2 S. dendroides |
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4 (2) |
Seed horizontal or oblique, ca. 2 mm in diam.; perianth appearing star-shaped in fruit. |
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3 S. glauca |
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Seed vertical, not exceeding 1.5 mm in diam.; perianth not star-shaped in fruit. |
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(5) |
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5 (4) |
Leaves filiform, semiterete, to 1 mm wide; seed sublustrous, without granular processes. |
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4 S. altissima |
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Leaves linear, slightly compressed, 1.5-3 mm wide; seed not lustrous, with granular processes. |
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(6) |
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6 (5) |
Flowers 3 or 4 per glomerule; perianth hemispheric to cupular, about as long as wide, 5-parted, segments slightly spreading at anthesis. |
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5 S. paradoxa |
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Flowers mostly solitary, rarely 2 or 3 per glomerule; perianth cylindric to obovoid, longer than wide, 5-lobed, segments remaining closed at anthesis. |
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6 S. linifolia |
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7 (1) |
Inflorescence ± a terminal panicle; perianth saccate inflated in fruit. |
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7 S. physophora |
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Inflorescence not a terminal panicle; perianth not inflated in fruit. |
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(8) |
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8 (7) |
Perianth segments distinctly veined; stigmas 3-5; seed extremely convex, scarcely pitted. |
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(9) |
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Perianth segments veinless; stigmas 2; seed slightly depressed, ± pitted. |
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(12) |
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9 (8) |
Perianth segments with a longitudinal, winglike keel abaxially. |
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(10) |
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Perianth segments without a keel abaxially. |
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(11) |
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10 (9) |
Leaf apex with a bristle; perianth segments keeled near apex, keel protruding beyond apex. |
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8 S. acuminata |
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Leaf apex without a bristle; perianth segments keeled throughout length. |
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9 S. pterantha |
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11 (9) |
Glomerules numerous flowered; stigmas 3, plumose; plants large, woody. |
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10 S. rigida |
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Glomerules 3-6-flowered; stigmas 3-5, not plumose; plants small, slender, herbaceous. |
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11 S. arcuata |
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12 (8) |
Leaves obovate, succulent, apex rounded or obtuse; glomerules mostly borne on dwarf, axillary branchlets. |
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(13) |
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Leaves not obovate, apex subobtuse or acute; glomerules all axillary. |
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(14) |
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13 (12) |
Perianth with a transverse, narrow, winglike process; seed distinctly pitted. |
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12 S. przewalskii |
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Perianth with a more developed transverse wing; seed obscurely pitted. |
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13 S. kossinskyi |
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14 (12) |
Seed obscurely pitted; anthers 0.3-0.5 mm. |
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(15) |
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Seed distinctly pitted; anthers ca. 0.2 mm. |
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(17) |
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15 (14) |
Shrubs small; leaf base articulate; stigmas erect, subulate. |
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18 S. australis |
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Herbs annual; leaf base not articulate; stigmas divaricate or slightly curved, filiform. |
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(16) |
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16 (15) |
Leaves usually sickle-shaped, apex obtuse. |
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19 S. crassifolia |
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Leaves straight or irregularly curved, apex acute. |
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20 S. salsa |
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17 (14) |
Perianth segments appearing unequally cornate. |
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14 S. corniculata |
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Perianth segments not as above. |
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(18) |
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18 (17) |
Leaf apex obtuse or acute, not mucronate; plants usually prostrate. |
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17 S. prostrata |
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Leaf apex usually distinctly mucronate; plants erect. |
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(19) |
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19 (18) |
Perianth segments each with a developed transverse wing, wings together appearing dish-shaped and 2.5-3.5 mm in diam. |
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15 S. heterophylla |
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Perianth segments each with a triangular, short, winglike process, processes together appearing star-shaped and not more than 2 mm in diam. |
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16 S. stellatiflora |
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List of lower taxa
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