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FOC | Family List | FOC Vol. 2-3 | Woodsiaceae

3. Woodsia R. Brown, Prodr. 158. 1810.

岩蕨属 yan jue shu

Plants small, usually epilithic. Rhizomes short, erect or ascending, rarely decumbent, scaly; scales lanceolate or linear-lanceolate, membranous. Fronds clustered; stipe shorter than lamina, articulate at various levels, i.e., from top to near base, or continuous in some species; lamina 1-pinnate or 1-pinnate-pinnatifid, lanceolate, herbaceous or subpapery, usually with articulate hairs or scales, rarely glabrous, base gradually tapering. Veins free, pinnate, not reaching laminar margin. Sori small, consisting of 3-18 sporangia, orbicular, on veinlets or at veinlet tips; indusia inferior, various, usually saucer- to cup-shaped, dissected into ciliate platelike segments or reduced into curly, multicellular hairs, sometimes exindusiate. Spores ellipsoid, monolete, perispore folded with granular, echinate or tuberculate ornamentations on surface. x = 39, 41.

About 38 species: widely distributed in N temperate regions and higher elevations of tropical America, Africa, and Madagascar; 20 species (eight endemic) in China.

The following taxon is excluded from the present treatment, pending further research: Woodsia shennongensis D. S. Jiang & D. M. Chen (J. Hunan Agric. Univ. 26(2): 88. 2000), described from Hubei (Shennongjia).

1 Stipe continuous, i.e., not articulate; indusia reduced into curly, multicellular hairs or apparently sori exindusiate (W. subg. Eriosorus (Ching) Shmakov)   (2)
+ Stipe articulate at various levels; indusia cup- or saucer-shaped, fimbriate, long ciliate   (9)
2 (1) Rhizome ascending or decumbent; lamina 1-imparipinnate, pinnae entire, slightly undulate, or crenate-lobed to pinnatifid   (3)
+ Rhizome erect, occasionally ascending; lamina 1-paripinnate or bipinnatifid, apical pinna absent, pinnae not entire; sori covered by curly, long multicellular hairs   (4)
3 (2) Pinnae entire or slightly undulate; sori exindusiate or with much-reduced indusia hidden.   19 W. cycloloba
+ Pinnae crenate-lobed to pinnatifid; indusia consisting of pale, long, crested hairs.   20 W. okamotoi
4 (2) Lamina 1-pinnate   (5)
+ Lamina bipinnatifid   (6)
5 (4) Rachis with sparse long hairs, occasionally with 1 or 2 scales; pinnae ovate, both surfaces sparsely shortly hairy.   13 W. macrospora
+ Rachis densely covered with scales; pinnae oblong-ovate, both surfaces densely covered with loose long hairs.   14 W. cinnamomea
6 (4) Plants 2-10 cm tall (usually ca. 5 cm tall); stipe and rachis chestnut or brown or brown- stramineous; lamina covered with hairs and/or scales abaxially; pinnae 4-10 pairs   (7)
+ Plants to 25 cm; stipes brown or brown-stramineous; lamina densely covered only with loose ferruginous hairs; pinnae 6-15 pairs   (8)
7 (6) Stipe and rachis chestnut; pinnae sessile; lamina covered with hairs and scales abaxially.   15 W. lanosa
+ Stipe and rachis brown or brown-stramineous; pinnae stalked to 1 mm; lamina covered with hairs only abaxially.   16 W. guizhouensis
8 (6) Pinnae ovate, basal pinnae not reduced or slightly reduced; indusia composed of glandular-tipped hairs.   17 W. andersonii
+ Pinnae elliptic, lower pinnae obviously reduced; indusia composed of ciliate platelike lobes.   18 W. rosthorniana
9 (1) Stipe articulate at or below middle (W. subg. Woodsia)   (10)
+ Stipe articulate at top, or sometimes at upper parts   (14)
10 (9) Lamina covered with long hairs and scales; stipes robust, castaneous or castaneous-brown   (11)
+ Lamina glabrous or with long hairs, without any scales; stipes slender, substramineous   (12)
11 (10) Lamina subglabrous, with sparse long hairs and scales on rachis; pinnae short, triangular-ovate or ovate, pinnatifid.   4 W. alpina
+ Lamina sparsely long hairy, rachis and costae scaly beneath; pinnae long, ovate-lanceolate, pinnatipartite nearly to costa.   5 W. ilvensis
12 (10) Lamina sparsely hairy abaxially, hairs long; pinnae shortly stalked.   3 W. shensiensis
+ Lamina glabrous; pinnae sessile   (13)
13 (12) Pinnae triangular-ovate, apex obtuse, 3-5 mm.   1 W. glabella
+ Pinnae subrhombic, apex acuminate or acute, up to 8 mm.   2 W. hancockii
14 (9) Indusia platelike; pinna base usually symmetrical or ?symmetrical (W. subg. Woodsia)   (15)
+ Indusia cup-shaped; pinna base asymmetrical or rarely symmetrical (W. subg. Acrolysis Nakai)   (16)
15 (14) Stipe 4-8 cm, brown or light castaneous; pinnae to 2 cm, pinnatifid.   11 W. subcordata
+ Stipe 1-3 cm, stramineous; pinnae 4-6 mm, pinnatifid into 3-5 segments.   12 W. sinica
16 (14) Pinnae below laminar apex free, elliptic-lanceolate or falcate-lanceolate; midvein scaly abaxially   (17)
+ Upper pinnae adnate, triangular-lanceolate or elliptic; midvein without scales or with few scales only abaxially   (18)
17 (16) Pinnae entire or undulate.   6 W. polystichoides
+ Pinnae pinnatilobate on both sides.   7 W. pilosa
18 (16) Only basal pinnae free, upper pinnae adnate, base symmetrical; rachis densely covered with long hairs, but without scales.   8 W. macrochlaena
+ Lower pinnae free, upper pinnae adnate, base asymmetrical; rachis covered with hairs and scales   (19)
19 (18) Pinnae sparsely hairy, herbaceous.   9 W. oblonga
+ Pinnae densely hairy on both surfaces, subpapery.   10 W. intermedia

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