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FOC | Family List | FOC Vol. 20-21 | Asteraceae

2. ARCTIUM Group

牛蒡群 niu bang qun

Authors: Zhu Shi & Werner Greuter

Herbs, biennial or perennial. Capitula homogamous, bisexual. Phyllaries ending in a straight, recurved, or hooked spine. Receptacle densely covered with long bristles. Florets bisexual. Corolla ± actinomorphic. Stamen filaments distinct, glabrous or papillose. Style branches distinct and divergent. Achene glabrous; apical rim ± inconspicuous, sometimes denticulate, not forming a crown; apical plate flat. Pappus of scabrid bristles not basally connate into a ring, falling off singly, or sometimes lacking.

About four genera and ca. 700 species: NW Africa, Asia, Europe; three genera and 14 species (two endemic) in China.

1 Outer and middle phyllaries ending in a hooked spine.   25 Arctium
+ Outer and middle phyllaries ending in a straight or recurved spine   (2)
2 (1) Capitula glabrous to cobwebby, when in clusters not surrounded by leaflike lanate bracts.   23 Cousinia
+ Capitula densely lanate, in clusters surrounded by leaflike lanate bracts.   24 Schmalhausenia


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