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2. ARCTIUM Group
牛蒡群 niu bang qun
Authors: Zhu Shi & Werner Greuter
Herbs, biennial or perennial. Capitula homogamous, bisexual. Phyllaries ending in a straight, recurved, or hooked spine. Receptacle densely covered with long bristles. Florets bisexual. Corolla ± actinomorphic. Stamen filaments distinct, glabrous or papillose. Style branches distinct and divergent. Achene glabrous; apical rim ± inconspicuous, sometimes denticulate, not forming a crown; apical plate flat. Pappus of scabrid bristles not basally connate into a ring, falling off singly, or sometimes lacking.
About four genera and ca. 700 species: NW Africa, Asia, Europe; three genera and 14 species (two endemic) in China.
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| 1 |
Outer and middle phyllaries ending in a hooked spine. |
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25 Arctium |
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Outer and middle phyllaries ending in a straight or recurved spine |
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(2) |
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| 2 (1) |
Capitula glabrous to cobwebby, when in clusters not surrounded by leaflike lanate bracts. |
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23 Cousinia |
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Capitula densely lanate, in clusters surrounded by leaflike lanate bracts. |
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24 Schmalhausenia |
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