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FOC | Family List | FOC Vol. 2-3 | Athyriaceae | Athyrium

52. Athyrium intermixtum Ching & P. S. Chui, Acta Bot. Boreal.-Occid. Sin. 6(1): 21. 1986.

中间蹄盖蕨 zhong jian ti gai jue

Rhizomes erect, apex densely clothed with castaneous or brown, narrowly lanceolate scales. Fronds caespitose; fertile fronds 70-85 cm; stipe brown-stramineous at base, upward shiny, 28-33 cm, ca. 3 mm in diam., base similarly scaly, upward glabrate; lamina 2-pinnate, ovate, 28-30 × 20-26 cm, base broadly cuneate, apex somewhat abruptly narrowed; pinnae ca. 13 pairs, basal 2(or 3) pairs of pinnae subopposite; upper pinnae alternate, ascending, stalked (stalk ca. 3 mm); lower 3 or 4 pairs of pinnae subequal, oblong-lanceolate, ca. 15 × 3-3.5 cm, base slightly narrowed, pinnate, apex shortly acuminate; upper pinnae gradually smaller; pinnules ca. 20 pairs, alternate, spreading, basal pinnules shortly stalked; upper pinnules sessile, oblong, ca. 1.8 cm × 6-8 mm, base broadly cuneate, auriculate on acroscopic side, pinnatilobate, apex obtuse-rounded; pinnule segments 4-6 pairs, with 2 or 3 teeth at margin, apex subrounded; veins somewhat visible adaxially, visible abaxially, 2- or 3-forked in segments. Lamina herbaceous when dried, dark brown, glabrate on both surfaces; rachis and costae abaxially brown-stramineous, with sparse short glandular hairs, with subulate spines on adaxial side at base of costa or costule. Sori elliptic, inframedial on acroscopic veins, 4-6 pairs per pinnule, biseriate on both sides of costule; indusia brown, elliptic, membranous, entire, persistent. Perispore surface without folds, with granular ornamentation.

● Roadsides on mountain slopes; 900-1600 m. Anhui, Zhejiang.


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