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FOC | Family List | FOC Vol. 2-3 | Athyriaceae | Athyrium

118. Athyrium spinulosum (Maximowicz) Milde, Bot. Zeitung (Berlin). 24: 376. 1866.

假冷蕨 jia leng jue

Cystopteris spinulosa Maximowicz, Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math., Seconde Pt. Sci. Nat. 9: 340. 1859; Asplenium spinulosum (Maximowicz) Miquel (1867), not (Blume) Mettenius (1859); Pseudocystopteris spinulosa (Maximowicz) Ching.

Rhizomes long creeping, ca. 2.5 mm in diam., blackish brown, with sparse broadly ovate membranous scales. Fronds distant; fertile fronds 45-50 cm; stipe blackish brown at base, upward stramineous, shiny, 22-26 cm, 2-3 mm in diam. at base, base clothed with broadly ovate to lanceolate scales, often densely yellow long pubescent, upper part glabrate, occasionally scaly; lamina triangular to ovate-triangular, 20-28 cm, nearly as broad as long or slightly broader, shortly acuminate or almost acute, 2- or 3-pinnate, with ca. 12 pairs of pinnae; basal pinnae longest, 17-20 × 6-8 cm at middle, elliptic to elliptic-oblanceolate, gradually narrowed toward base, subequilateral at base, with stalk 5-8 mm, acuminate, subopposite, 2-3.5 cm apart from upper pinnae, ascending; basal pinnae with pinnules 12-14-paired, anadromous, ascending or spreading, oblong-lanceolate to ovate-triangular; lower pinnules gradually shorter, obtuse or acuminate, equilateral, rounded or rounded-cuneate at base, sessile or slightly adnate to costae, rarely very shortly stalked, with long pointed teeth at margin; middle pinnules longest, 4-5 cm × 8-15 mm; secondary pinnules 10 pairs or fewer, ascending, oblong or narrowly ovate, 5-8 × 2.5-3.5 mm, usually decurrent to costular wings, sharply lobed or with thin, long, sharp teeth; segments sharply toothed; second basal and upper pinnae lanceolate, narrower, subopposite or alternate, not narrowed toward base, shortly stalked or sessile; basal pair of pinnules catadromous or subopposite; pinnule segments with pointed, long, sharp teeth. Veins visible on surfaces, pinnate, simple, terminating in tooth tip. Lamina herbaceous when dried, green or greenish brown, costule and midrib bases with some short spinulose processes on adaxial surface, rachis, and costae with sparse short hairs adaxially, glabrate abaxially. Sori orbicular, abaxial on veins, 1 to several pairs per every ultimate segment; indusia brownish, almost orbicular-reniform, membranous. Perispore thin, hyaline, folded. 2n = 80.

Damp areas in coniferous and mixed forests, in shrubs, bamboo groves; 800-3000 m. Heilongjiang, Henan, Jilin, Liaoning, Nei Mongol, Shaanxi, Sichuan [Japan, Korea, E Russia].


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