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FOC | Family List | FOC Vol. 2-3 | Athyriaceae | Athyrium

119. Athyrium subtriangulare (Hooker) Beddome, Ferns Brit. India. t. 242. 1867.

三角叶假冷蕨 san jiao ye jia leng jue

Asplenium subtriangulare Hooker in Hooker & Baker, Syn. Fil. 225. 1867; A. spinulosum (Blume) Mettenius var. subtriangulare (Hooker) C. B. Clarke; Athyrium hookerianum T. Moore; A. longipes Christ; A. sikkimense (Bir) Á. Löve & D. Löve; A. spinulosum (Maximowicz) Milde var. subtriangulare (Hooker) C. Christensen; A. subtriangulare var. sikkimense Bir; Pseudocystopteris decipiens Ching & S. K. Wu; P. longipes (Christ) Ching; P. purpurascens Ching & S. K. Wu; P. reflexipinnula Ching & S. K. Wu; P. remota Ching; P. sikkimensis (Bir) Ching; P. sparsa Ching & S. K. Wu; P. spinulosa (Maximowicz) Ching var. taipaishanensis Ching; P. subtriangularis (Hooker) Ching; P. tibetica Ching.

Rhizomes long creeping, 3-4 mm in diam., blackish brown, sparsely scaly; scales brownish, ovate-lanceolate. Fronds distant; fertile fronds (13-)60-70(-90) cm; stipe brownish at base, upward stramineous, (5-)30-40(-55) cm, (1-)3-4(-5) mm in diam. at base, base densely scaly, often densely long yellow pubescent, upper part occasionally with brownish lanceolate membranous scales; lamina 2- or 3-pinnate, broadly ovate-triangular to ovate-triangular, (8-)25-35(-40) × (10-)20-30(-45) cm, acuminate; pinnae (10-)15-20 pairs, subalternate, ascending, approximate; basal pinnae largest, 14-20(-28) × 5-9(-12) cm at middle, elliptic to elliptic-oblanceolate, acuminate, gradually narrowed and subequilateral at base, with stalks 5-8 mm, subopposite, (1-)4-5 cm apart from upper pinnae; pinnules of basal pinnae anadromous, (10-)14-20 pairs, spreading or slightly ascending, upper pinnules gradually shortened to base, oblong-lanceolate or ovate-triangular, obtuse, usually with patulous teeth; pinnules of middle pinnae oblong-lanceolate, 2.5-4.5(-7) × 1-1.5 cm at base, shortly acuminate or acute, sessile, subtruncate, separate or partly adnate to costae, approximate; secondary pinnules 10-12 pairs, spreading or slightly ascending, oblong, 5-8 × 2.5-4 mm, obtuse, decurrent to costular wings, dissected or with patulous teeth; second basal pinnae similar to but slightly smaller than basal pinnae, oblong; upward pinnae lanceolate, acuminate, not narrowed toward base; ultimate segments serrate. Veins visible on surfaces, pinnate in ultimate pinnule, simple, terminating into teeth. Lamina herbaceous when dried, green or brown-green, costules and midribs with very short spines on adaxial surface, rachis, costae, and costules ± shortly hairy, veins with sparse glandular short hairs or multicellular hairs abaxially, costa base occasionally with few small scales. Sori orbicular, abaxial on veins, 1 to several pairs per ultimate segment; indusia brownish, membranous, orbicular-reniform. Perispore thin, hyaline, folded. 2n = 80*, 160.

Damp areas in forests; (500-)2000-4100 m. Chongqing, Gansu, Hubei, Qinghai, Shaanxi, Sichuan, SE Xizang, Yunnan [Bhutan, N India, Nepal].


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