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FOC | Family List | FOC Vol. 23 | Cyperaceae

11. CAREX sect. RACEMOSAE G. Don in Loudon, Hort. Brit. 376. 1830.

褐穗薹草组 he sui tai cao zu

Authors: Prof. Song-Yun Liang & Tetsuo Koyama

Involucral bracts leaflike or setaceous. Spikes erect, rarely pendent; terminal spike gynaecandrous, rarely male; lateral spikes female, sometimes with a few apical male flowers. Female glumes dark purple-brown. Utricles yellow-green or pale brown, usually tinged with dark purple-brown, elliptic, ovate, or obovate, membranous or leathery, glabrous, minutely papillose or punctate, apex shortly beaked, orifice entire, emarginate, or 2-toothed. Style base thickened; stigmas 3, rarely 2.

About 60 species: Asia, Europe, North America; 28 species (ten endemic) in China.


1 Female glumes 9-10 mm.   37 C. praeclara
+ Female glumes less than 8 mm   (2)
       
2 (1) Utricles densely papillose, gray-green   (3)
+ Utricles smooth or minutely punctate, yellowish green, stramineous, pale brown, or dark purple-red   (5)
       
3 (2) Female glumes acuminate or with short awn at apex, longer than utricles.   38 C. tarunensis
+ Female glumes obtuse at apex, equaling or slightly shorter than utricles   (4)
       
4 (3) Female spikes 5-12 mm, sessile.   39 C. meyeriana
+ Female spikes 10-25 mm, shortly stipitate.   40 C. hongyuanensis
       
5 (2) Upper 3 or 4 spikes male, rarely with a few female flowers at base.   41 C. polymascula
+ Terminal spikes male or gynaecandrous or all spikes gynaecandrous   (6)
       
6 (5) All spikes gynaecandrous.   42 C. aristulifera
+ Spikes gynaecandrous or with terminal spike male   (7)
       
7 (6) Terminal spike male (rarely bisexual in C. melanantha)   (8)
+ Terminal spike gynaecandrous   (11)
       
8 (7) Female glumes broadly white hyaline on margins.   45 C. moorcroftii
+ Female glumes narrowly white hyaline on margins or not   (9)
       
9 (8) Utricles 2-toothed at orifice.   46 C. minxianensis
+ Utricles emarginate at orifice   (10)
       
10 (9) Spikes distant; leaves soft.   43 C. augustinowiczii
+ Spikes approximate at culm apex, only lowest spike distant; leaves slightly stiff.   44 C. melanantha
       
11 (7) Utricles scabrous above on margins.   47 C. obscura
+ Utricles smooth on margins   (12)
       
12 (11) Female glumes with a scabrous long awn at apex.   48 C. gmelinii
+ Female glumes without scabrous long awn at apex   (13)
       
13 (12) Utricles veined   (14)
+ Utricles nerveless (except for 2 costas on edges)   (19)
       
14 (13) Utricles elliptic or elliptic-lanceolate   (15)
+ Utricles obovate-elliptic, obovate, or oblanceolate   (16)
       
15 (14) Spikes distant; female glumes pale or light ferruginous; orifice of utricles yellow-green, 2-toothed.   59 C. peiktusani
+ Spikes approximate; female glumes dark purple; orifice of utricles purple-red, emarginate.   60 C. schneideri
       
16 (14) Utricles inflated   (17)
+ Utricles not inflated   (18)
       
17 (16) Utricles 2-2.2 mm; female glumes broadly ovate, ca. 1.2 mm.   61 C. lehmannii
+ Utricles (2.5-)3-3.3 mm; female glumes oblong, 2-2.5 mm.   62 C. montis-wutaii
       
18 (16) Utricles oblanceolate or narrowly elliptic, beak 2-toothed at orifice.   63 C. serreana
+ Utricles obovate or elliptic-obovate, beak short and subtruncate at orifice.   64 C. caucasica
       
19 (13) Spikes contiguous forming a capitate or spicate inflorescence   (20)
+ Spikes loosely disposed in racemose inflorescence   (22)
       
20 (19) Utricles membranous, vesicular, nerveless.   51 C. melanocephala
+ Utricles leathery, granulose or glabrous, 3-5-veined   (21)
       
21 (20) Rhizome long; utricles ovate, glabrous, 4-5 mm; sandy plants.   49 C. sabulosa
+ Rhizome short, culms loosely tufted; utricles obovate, minutely granulose, 2.5-3 mm.   50 C. melananthiformis
       
22 (19) Female glumes 2-3 mm; female spikes 4-15 mm   (23)
+ Female glumes 4-6 mm; female spikes (20-)25-35 mm   (25)
       
23 (22) Utricles inflated, horizontally patent at maturity.   57 C. hancockiana
+ Utricles compressed or slightly inflated, not horizontally patent at maturity   (24)
       
24 (23) Culms 30-60 cm tall; female glumes white hyaline on margins.   55 C. angarae
+ Culms 15-25 cm tall; female glumes not white hyaline on margins.   56 C. duthiei
       
25 (22) Utricles stramineous, sometimes upper yellow-brown or purple punctate, compressed.   52 C. kansuensis
+ Utricles dark brown to purple-brown   (26)
       
26 (25) Female glumes 6-6.5 mm; nutlets ca. 2.5 mm.   53 C. bijiangensis
+ Female glumes 3.5-4 mm; nutlets 1.5-1.7 mm   (27)
       
27 (26) Utricles elliptic or ovate, shallowly 2-toothed at orifice.   58 C. atrata
+ Utricles narrowly oblong, obliquely truncate at orifice.   54 C. obliquitruncata

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