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FOC | Family List | FOC Vol. 8 | Crassulaceae | Sedum

3. Sedum sect. Sedum

景天组 jing tian zu

Leaf base sessile or narrowed into a petiole, spurred. Inflorescence branches often scorpioid, rarely corymbiform. Flowers mostly sessile, basal flower sometimes pedicellate. Sepals spurred or spurless. Petals yellow. Carpels mostly subdivergent to spreading stellately at maturity, sometimes erect, adaxially gibbous, base connate. Follicles usually spreading-divergent, sometimes erect, adaxially gibbous. Seeds mostly papillate, sometimes smooth.

More than 60 species: Asia, Europe; 49 species (34 endemic) in China.


1 Plants erect, rarely prostrate; leaves usually spurless (shortly spurred in S. leptophyllum).   (2)
+ Plants subprostrate, ascending, or decumbent; leaves usually spurred; sepals spurred, rarely spurless.   (8)
       
2 (1) Pseudopetiole conspicuous.   (3)
+ Pseudopetiole inconspicuous, leaves tapered toward base.   (5)
       
3 (2) Carpels 4; follicles many seeded   75 S. tetractinum
+ Carpels 5; follicles 1-3-seeded.   (4)
       
4 (3) Bracts orbicular to suborbicular; leaves rhombic-elliptic, 3-6 cm, apex attenuate or retuse to emarginate   73 S. oligospermum
+ Bracts linear-spatulate; leaves ovate to suborbicular, 1.5-3 cm, apex rounded   74 S. wilsonii
       
5 (2) Carpels and nectar scales 3; seeds few.   (6)
+ Carpels and nectar scales 4; seeds numerous.   (7)
       
6 (5) Leaves verticillate or opposite, spatulate-oblong   76 S. triactina
+ Leaves alternate, rarely verticillate, oblanceolate to linear-lanceolate   77 S. leptophyllum
       
7 (5) Leaves narrowly oblanceolate, 1-2.5 cm; sepals triangular to narrowly elliptic; petals 4-4.5 mm, apex shortly mucronate   78 S. hakonense
+ Leaves oblanceolate, 2.5-4.5 cm; sepals narrowly triangular; petals ca. 7 mm, apex acute   79 S. dongzhiense
       
8 (1) Rootstock with root shoots or white, globose latent bulbs.   (9)
+ Rootstock absent, roots slender.   (12)
       
9 (8) Rootstock ascending, root shoots present.   (10)
+ Rootstock slender, white, globose latent bulbs present.   (11)
       
10 (9) Plants 5-20 cm tall; leaves spatulate to broadly obovate, 12-20 mm   80 S. leucocarpum
+ Plants ca. 7 cm tall; leaves ovate, 3-4 mm   81 S. wenchuanense
       
11 (9) Leaves alternate, 3-15 × ca. 2 mm; sepals 1.5-7 mm   82 S. tianmushanense
+ Leaves opposite at base of stem, 4-verticillate at middle, alternate at apex, 12-30 × 1-2 mm; sepals 2-2.5 mm   83 S. latentibulbosum
       
12 (8) Sepals equaling or longer than petals or, if shorter than the petals, all sepals equal.   (13)
+ Sepals shorter than petals or, if longer than petals, sepals unequal.   (14)
       
13 (12) Plants annual; petals triangular, apex acute   84 S. woronowii
+ Plants perennial; petals oblong-ovate, apex long mucronate   85 S. multicaule
       
14 (12) Plants suffrutescent, tufted; sepals 1/4-1/3 as long as petals   86 S. polytrichoides
+ Plants mostly not tufted, leathery; sepals to 1/2 as long as petals.   (15)
       
15 (14) Leaves usually verticillate (sometimes opposite or less often alternate in S. erythrospermum).   (16)
+ Leaves usually alternate or opposite (verticillate or verticillate and alternate in S. cryptomerioides, S. dimorphophyllum, and S. subtile).   (24)
       
16 (15) Leaves usually 1.5-3 cm.   (17)
+ Leaves 0.5-1.4 cm.   (19)
       
17 (16) Leaves oblanceolate to oblong   117 S. sarmentosum
+ Leaves linear to linear-oblanceolate.   (18)
       
18 (17) Petal apex not mucronate   118 S. lineare
+ Petal apex long mucronate   119 S. grammophyllum
       
19 (16) Leaves opposite or 3- or 4-verticillate.   (20)
+ Leaves 4- or 5-verticillate.   (22)
       
20 (19) Plants annual; sterile stems absent   113 S. erythrospermum
+ Plants perennial; sterile stems present.   (21)
       
21 (20) Leaves 3-verticillate and linear on flowering stems, 3-verticillate and obovate on sterile stems; sepals linear-obovate   111 S. jiulungshanense
+ Leaves on flowering stems and sterile stems 3- or 4-verticillate, broadly linear to lanceolate; sepals broadly linear to suboblong   112 S. onychopetalum
       
22 (19) Leaves 4- or 5-verticillate, differently shaped basally and apically on stem   116 S. kiangnanense
+ Leaves 4-verticillate, similarly shaped basally and apically on stem.   (23)
       
23 (22) Leaf apex acute; nectar scales spatulate-oblong; anthers blackish   114 S. phyllanthum
+ Leaf apex obtuse; nectar scales oblong-cuneate; anthers not blackish   115 S. yvesii
       
24 (15) Leaves verticillate and alternate apically on stems, verticillate basally on old stems   87 S. dimorphophyllum
+ Leaves alternate or opposite (verticillate apically on stems in S. cryptomerioides and occasionally in S. subtile).   (25)
       
25 (24) Leaves usually conspicuously pseudopetiolate.   (26)
+ Leaves inconspicuously pseudopetiolate.   (27)
       
26 (25) Leaf base spurred; sepals equal; bracts very large, resembling stem leaves   120 S. concarpum
+ Leaf base spurless; sepals unequal; bracts very small, mostly obscure   121 S. engleri
       
27 (25) Leaves opposite (sometimes alternate or less often verticillate in S. actinocarpum).   (28)
+ Leaves alternate (opposite basally on stems in S. bulbiferum and occasionally in S. formosanum).   (31)
       
28 (27) Stems 3-7 cm   110 S. baileyi
+ Stems 10-25 cm.   (29)
       
29 (28) Leaf apex emarginate   108 S. emarginatum
+ Leaf apex truncate.   (30)
       
30 (29) Stems erect; carpels suboblong to ovoid, ca. 2.5 mm; follicles horizontally spreading   107 S. actinocarpum
+ Stems basally prostrate; carpels lanceolate, ca. 5 mm; follicles obliquely spreading   109 S. makinoi
       
31 (27) Plants pubescent throughout.   (32)
+ Plants glabrous throughout.   (33)
       
32 (31) Leaves obovate; sepals ovate   88 S. giajae
+ Leaves spatulate; sepals spatulate   89 S. lungtsuanense
       
33 (31) Bulbils usually present in leaf axils   90 S. bulbiferum
+ Bulbils absent in leaf axils.   (34)
       
34 (33) Sterile stems absent.   (35)
+ Sterile stems present.   (41)
       
35 (34) Sepals wider at base than at middle.   (36)
+ Sepals narrower at base than at middle.   (37)
       
36 (35) Sepals triangular   100 S. microsepalum
+ Sepals linear-ovate   101 S. hangzhouense
       
37 (35) Leaves 0.6-1 cm; seeds minutely papillate or reticulate.   (38)
+ Leaves 1.2-3 cm; seeds smooth.   (39)
       
38 (37) Leaves ca. 6 × 1.5 mm; seeds reticulate   102 S. morrisonense
+ Leaves 6-10 × 3-5 mm; seeds papillate   103 S. nokoense
       
39 (37) Flowering stems branched from base   104 S. formosanum
+ Flowering stems simple, rarely apically branched.   (40)
       
40 (39) Leaves oblanceolate to narrowly elliptic-lanceolate; sepals 1.5-3 × ca. 0.5 mm; seeds yellow   105 S. parvisepalum
+ Leaves linear-cuneate, spatulate, or obovate; sepals 3-5 × 1-1.5 mm; seeds brown   106 S. alfredii
       
41 (34) Leaves of flowering stems terete or ± flat.   (42)
+ Leaves of flowering stems flat.   (43)
       
42 (41) Leaves of flowering stems very crowded, 0.3-0.5 cm; sepals to 3 mm   91 S. uniflorum
+ Leaves of flowering stems ± remote, 7-10 cm; sepals to 4 mm   92 S. japonicum
       
43 (41) Leaves of flowering stems minutely papillate on distal margin; sepals minutely papillate; follicles stellately subhorizontally spreading   93 S. planifolium
+ Leaves of flowering stems not minutely papillate on margin; sepals not minutely papillate; follicles stellately divaricate but not horizontally spreading.   (44)
       
44 (43) Sepals spurless; leaves of sterile stems rosulate   95 S. sekiteiense
+ Sepals spurred; leaves of sterile stems not rosulate.   (45)
       
45 (44) Leaves opposite or 3-6-verticillate basally on stem or verticillate and crowded apically on stem.   (46)
+ Leaves all alternate.   (47)
       
46 (45) Leaves opposite or 3-6-verticillate basally on stem   96 S. subtile
+ Leaves verticillate and crowded apically on stem   99 S. cryptomerioides
       
47 (45) Sepals subequal; seeds smooth   94 S. hengduanense
+ Sepals unequal; seeds papillate.   (48)
       
48 (47) Petals narrowly elliptic-lanceolate   97 S. tosaense
+ Petals oblong-lanceolate   98 S. nanchuanense

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