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FOC | Family List | FOC Vol. 10 | Fabaceae | Fabaceae Tribe GALEGEAE

45. Astragalus sect. Helmia Bunge, Mém. Acad. Imp. Sci. Saint Pétersbourg, sér. 7. 11(16): 111. 1868.

乌拉尔组 wu la er zu

Plants often dwarf, acaulescent or nearly so, caespitose, covered with bifurcate hairs. Stipules free or adnate to petiole, vaginate-connate behind stem. Racemes shortly to long pedunculate, rarely subsessile, few or up to 20-flowered; bracts whitish hyaline. Flow­ers with a short pedicel or subsessile. Calyx campanulate to tubular. Petals yellow or purple. Legumes sessile to subsessile, com­pressed laterally or inflated, keeled ventrally, narrowly grooved to flattened dorsally, 1-locular to 2-locular; valves thinly leathery, densely covered with mostly spreading hairs.

Twenty-four species: China, India, Kashmir, Mongolia, Nepal, Pakistan; C Asia, E Europe to Siberia; 16 species (ten endemic) in China.

1 Leaves simple, 10-50 × ca. 1 mm.   298 A. efoliolatus
+ Leaflets in at least 1 pair but mostly in more   (2)
2 (1) Leaflets in 10-13 pairs, villous with spreading hairs; bracts 4-5 mm; petals yellow.   304 A. pendulatopetalus
+ Leaflets in 1-6 pairs, covered with ± appressed hairs; bracts shorter; petals mostly purple to violet   (3)
3 (2) Standard up to 10(-11) mm   (4)
+ Standard distinctly longer   (8)
4 (3) Leaflets linear, 8-10 × as long as wide   (5)
+ Leaflets narrowly elliptic, up to 5 × as long as wide   (6)
5 (4) Plants 3-12 cm tall, at base with numerous non-flowering off-shoots; calyx campanulate, 4-6 mm; wing limbs retuse to slightly emarginate.   303 A. nematodioides
+ Plants without non-flowering off-shoots; calyx tubular; wing limbs distinctly emarginate.   306 A. vladimirii-komarovii
6 (4) Leaflets in 4 or 5 pairs, 6-10 × 1-2 mm; legumes 2-locular, subacute, without beak.   296 A. divnogorskajae
+ Leaflets in 1-3 pairs, 1-6 × 0.5-1 mm; legumes 1-locular or incompletely 2-locular, with a beak 1-2 mm   (7)
7 (6) Stipules 1.5-2 mm, white and black hairy; standard ca. 10 mm; peduncles 4-9 cm; legumes ca. 1.5 mm high and ca. 1 mm wide, subappressed hairy, incompletely 2-locular.   297 A. dshangartensis
+ Stipules 3-5 mm, white hairy; standard 6-8 mm; peduncles 1-2.5 cm; legumes inflated, with ± spreading hairs, nearly 1-locular.   291 A. arnoldii
8 (3) Leaflets 10-30(-40) mm; stipules 4-12 mm, adnate to petiole for 3-4 mm   (9)
+ Leaflets up to 11(-15) × ca. 3 mm; stipules up to 5 mm, free from petiole or shortly adnate to it   (11)
9 (8) Calyx 7-10 mm, widely campanulate, base slightly inflated ovoid, urn-shaped, only white hairy; standard 15-20 × 6-7 mm, obovate, below middle slightly constricted.   294 A. cupulicalycinus
+ Calyx at least 10 mm, tubular, at least with some black hairs; standard at least 20 mm but mostly distinctly longer, not constricted   (10)
10 (9) Only stipules of lower leaves connate, those of upper leaves free from each other; leaflets with hairs all similar, 0.5-1 mm; standard 20-23 mm, ovate with rather long claw; legumes covered with extremely asymmetrically bifurcate, spreading, flexuous hairs ca. 1 mm, 1-locular.   292 A. burtschumensis
+ All stipules high connate behind stem; leaflets with hairs 0.5-0.8 mm and with distinctly thicker hairs 1-1.8 mm; standard 21-30 mm, obovate with short claw; legumes unknown.   302 A. eerqisiensis
11 (8) Stipules 1-3 mm; standard up to 17 mm, elliptic or obovate, never constricted; legumes (ovary) appressed hairy   (12)
+ Stipules 2.5-5(-6) mm; standard 17-24(-26) mm, in middle slightly constricted or not; legumes villous, with spreading hairs   (13)
12 (11) Stipules 1-2 mm, nearly free from petiole, behind stem shortly connate; leaflets in 2-5 pairs; bracts ca. 1 mm, white and black hairy; calyx covered with subappressed, flexuous, black hairs with some longer, spreading, white and black hairs; legumes (immature) linear, falcate, ca. 15 mm.   305 A. persimilis
+ Stipules 2-3 mm, adnate to petiole for ca. 1 mm, behind stem connate to top; leaflets in 1 pair; bracts 1.5-3 mm, predominantly white hairy; calyx densely covered with appressed white and black hairs; legumes unknown.   300 A. hoshanbaoensis
13 (11) Stipules free from petiole; calyx 7-8 mm; standard narrowly elliptic-pandurate, ca. 6 mm wide, constricted in middle; wing limbs obliquely incised; legumes (6-)8-10 mm, nearly 1-locular.   293 A. chomutowii
+ Stipules shortly adnate to petiole; calyx at least 8 mm but mostly distinctly longer and up to 12 mm; standard 8-10 mm wide, if only ca. 6 mm wide (in A. kasachstanicus) then oblong, not constricted, and wing limbs obtuse; legumes 18-23 mm, 2-locular   (14)
14 (13) Standard oblong, ca. 6 mm wide; wing limbs obtuse; legumes with tangled, flexuous, spreading hairs ca. 3 mm, sitting on small tubercles.   301 A. kasachstanicus
+ Standard obovate, 8-10 mm wide; wing limbs retuse or incised; hairs of legumes not sitting on small tubercles   (15)
15 (14) Petals pale yellowish or white; standard obovate, not constricted; wing limbs slightly retuse; legumes with a stipe 1-2 mm.   295 A. depauperatus
+ Petals pale purplish; standard constricted in middle; wing limbs distinctly emarginate; legumes sessile.   299 A. heptapotamicus


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