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FOC | Family List | FOC Vol. 10 | Fabaceae

25. Tribe GALEGEAE

山羊豆族 shan yang dou zu

Authors: Langran Xu, Xiangyun Zhu, Bojian Bao, Mingli Zhang, Hang Sun, Dietrich Podlech, Stanley L. Welsh, Hiroyoshi Ohashi, Kai Larsen & Anthony R. Brach

Herbs or shrubs, with simple or T-shaped hairs; glands or glandular punctae sometimes present. Leaves epulvinate or pulvinus reduced, imparipinnate or paripinnate, with many opposite to irregularly arranged or rarely conjugate leaflets, rarely 1-3-foliolate; stipules free or adnate to petiole, estipellate. Flowers in axillary racemes, spikes, or rarely solitary. Calyx campanulate to tubular; standard clawed or narrowed to base; wings auriculate; keel blunt to apiculate. Stamens diadelphous, rarely monadelphous; anthers usually uniform, but slightly dimorphic and with confluent thecae in Glycyrrhiza. Ovary few to many ovuled (sometimes 1-seeded); style slender, bearded or not, with a terminal or lateral stigma. Legumes compressed, angled or inflated, sometimes with sutured margins intruded or longitudinally septate, occasionally torulose, dehiscent or indehiscent. Seeds oblong-reniform, estrophiolate.

About 24 genera and 2900-3200 species: principally in Asia, Europe, and North America, but extending thinly in mountainous and/or drier places to S Africa, Australia, and temperate South America; 11 genera and 586 species (324 endemic, two introduced) in China.

Galega officinalis Linnaeus (Sp. Pl. 2: 714. 1753), probably native to SW Asia (Caucasus), is cultivated in China.

1 Style bearded, sometimes just a tuft of hairs below stigma on one side; wings and keel never interlocking (subtribe Coluteinae)   (2)
+ Style glabrous, but stigma sometimes penicillate (sometimes inner side of style upper part hairy in Astragalus); wings and keel mostly interlocking with bosses on keel and matching sockets on wings (indistinct only in Gueldenstaedtia and Tibetia)   (5)
2 (1) Leaves reduced to scales; flowers solitary; legumes compressed.   145 Eremosparton
+ Leaves imparipinnate, 7-25-foliolate; flowers in racemes; legumes inflated   (3)
3 (2) Flowers yellow; standard with 2 callosities above claw; shrubs.   143 Colutea
+ Flowers red to violet, rarely yellow; standard without callosities above claw; shrubs or herbs   (4)
4 (3) Plants with medifixed hairs; legumes inflated subglobose.   144 Sphaerophysa
+ Plants with basifixed hairs; legumes narrowly ellipsoid to ovoid.   138 Phyllolobium
5 (1) Anther thecae confluent toward apex; leaves pulvinate, spirally arranged; some hairs glandular-peltate or -capitate (subtribe Glycyrrhizinae).   148 Glycyrrhiza
+ Anther thecae separate; leaf pulvinus reduced or absent; plants without peltate glandular hairs   (6)
6 (6) Legume nerves oblique, closely parallel, raised at maturity, confluent toward placental margin; vexillary filament connate with others; keel petals not auriculate; leaflet nerves extending to margin (subtribe Galeginae).  

Galega (see note above)

+ Legume nerves transverse, confluent over middle part of valves, fine or inconspicuous; vexillary filament generally free; keel petals generally auriculate; leaflet nerves usually looped within margin or inconspicuous (subtribe Astragalinae)   (7)
7 (6) Calyx base usually oblique, upper side ± slightly gibbous; wings pinnately nerved; legume valves generally twisting on dehiscing   (8)
+ Calyx base symmetric or suboblique; wings usually palmately nerved; legume valves not twisting   (9)
8 (7) Stem obvious; stipules membranous, separated from petiole; calyx campanulate.   142 Chesniella
+ Stem abbreviated, plants appearing acaulescent; stipules herbaceous, adnate to petiole; calyx tubular.   141 Chesneya
9 (7) Keel petals half as long as wings; style shorter than or as long as ovary   (10)
+ Keel petals subequal to or slightly shorter than wings; style longer than ovary   (11)
10 (9) Style incurved; upper 2 calyx lobes free; seeds scrobiculate; stipules free and adnate to base of petiole.   146 Gueldenstaedtia
+ Style curved in right angle; upper 2 calyx lobes connate; seeds smooth, marbled-spotted; stipules connate below apex and opposite to leaves.   147 Tibetia
11 (9) Keel petals obtuse; legume 1-locular or with a septum intruding from abaxial suture; leaflets ± symmetric.   139 Astragalus
+ Keel petals apiculate; legume 1-locular or with a septum intruding from adaxial suture; leaflets basally ± oblique or, if narrow, falcately incurved.   140 Oxytropis

  • List of lower taxa


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