Plants medium-sized, terrestrial. Rhizomes short and erect or ascending, including stipe bases with sparse scales; scales lanceolate, dark brown, thick, margins setaceous. Fronds clustered; stipes grayish stramineous, grooved adaxially, throughout with hairs; laminae elliptic or triangular, cordate at bases, 1-pinnate or pinnatifid, or simple, acuminate at apices; laminae if pinnate then with 1-6 pairs of lateral pinnae, pinnae broadly lanceolate, bases rounded, symmetrical, margins entire, free or adnate to rachises, obliquely spreading, apices acuminate; costae raised on both sides, veinlets evident, oblique distally and reaching margins, reticulate, thick and evident, areoles in 3 or 4 rows, slightly tetragonal or pentagonal, without included veinlet or simple or forked included veinlets. Laminae papery, dark brown when dry, rough, with dense hooked thick hairs on both surfaces. Sori scattered on included veinlets, exindusiate; sporangia bearing erect acicular hairs near annuli; spores elliptic, echinate. x = 36.

Four species: Bhutan, China, N India, Japan, Myanmar, Nepal, Thailand, Vietnam; four species (two endemic) in China.

(Authors: Lin Youxing (林尤兴); Kunio Iwatsuki)