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Pakistan | Family List | Pakistan V. 204 | Chenopodiaceae

5. Atriplex Linnaeus, Sp. Pl. 1052. 1753; Gen. Pl. ed. 5: 472. 1754; Hook.f., Fl. Brit. Ind. 5: 6. 1886; Kom., Fl. URSS 6: 75. 1936; Aellen in Tutin et al., Fl. Europ. 1: 95. 1964; in Davis, Fl. Turk. 2: 305. 1966; Hedge in Rech. f., Fl. Iran. 172: 63. 1997.

Herbs or shrubs, farinose or glabrous, often whitish, with usually flat, opposite or alternate leaves, monoecious rarely dioecious. Flowers polygamous, usually unisexual, in terminal or axillary clusters, forming a spike-like or paniculate inflorescence, male flowers with usually 5 perianth segments and 5 stamens; pistillate flowers usually without perianth but usually enclosed within 2, free or partly united bracteoles (valves), rarely with 4-5-lobed perianth segments then bracteoles absent. Styles and stigmas 2. Utricle usually enclosed within bracteoles (valves), pericarp membranous, mostly free or almost so; seed usually vertical, rarely horizontal or pendulous (especially in those with perianth), often dimorphic, smooth.

About 200 species, mainly distributed in deserts or saline habitats; 11 species are reported from Pakistan.

A difficult genus with some species showing great variability in the form of the bracteoles in fruit.

1 Fruiting valves free to the base; annual herbs.   (2)
+ Fruiting valves ± united at least at the base; annuals, biennials or perennials.   (4)
2 (1) Female flowers dimorphic, some with a perianth and fruit with a horizontal seed, some without a perianth and fruit with a vertical seed.   (3)
+ Female flowers monomorphic, without a fruit with a vertical seed.(Fig.11, A-C1).   12 A. dimorphostegia
3 (2) Leaves with a dense vesicular scaly indumentum on lower side and almost glabrous above, bicolorous.   8 A. aucheri
+ Leaves glabrous or almost so on both sides, concolorous. (Fig.12, G).   7 A. hortensis
4 (1) Fruiting valves 4-winged; dioecious shrubs. (Fig.9, A-G).   4 A. canescens
+ Fruiting valves not 4-winged; mono-ecious herbs or shrubs.   (5)
5 (4) Fruiting valves tuberculate.   (6)
+ Fruiting valves not tuberculate   (7)
6 (5) Upper leaf surface glabrous; petiole on middle cauline leaves 15-20 mm. (Fig.12, A-F).   10 A. schugnanica
+ Upper leaf surface with an indumentum; petiole on middle cauline leaves less than 10 mm. (Fig.10, A-D1)   6 A. tatarica
7 (5) Shrubs with a sub-orbicular to reniform, ± flat fruit; valves fused only at the base; leaves entire or obsoletely repand-lobed.   (8)
+ Shrubs, perennial or annual herbs with variable fruits; valves clearly fused; leaves entire or clearly serrate.   (10)
8 (7) Stem short, 10-80 cm; glomerules few to many-flowered; fruiting spikes ± lax; fruit valves 5-10 x 5-2 mm.   (9)
+ Stem long, 2-3 m; glomerules many-flowered forming dense lateral and terminal spikes; fruiting spike ± leafless; dense fruit valves c. 4-5 mm, thick-textured.   11 A. halimus
9 (8) Fruit valves 5-6 x 4-6 mm; leaves oblong-obovate, attenuate into a 0.5-1.5 mm petiole. (Fig.8, G-I).   2 A. stocksii
+ Fruit valves 7-10 x 10-12 mm; leaves rhomboid to suborbicular, broadly cuneate to subcordate with a 2-10 mm petiole on lower leaves. (Fig.8, G-I)   3 A. griffithii
10 (7) Annual herbs; leaves not thick, entire to serrate.   (11)
+ Perennial shrub; leaves thick-textured, entire to sinuate dentate. (Fig.8, A-F1)   1 A. leucoclada
11 (10) Inflorescence with at least some distinct vesicular hairs.   5 A. pamirica
+ Inflorescence without distinct vesicular hairs, glabrescent. (Fig.11, D-F1)   9 A. lasiantha

  • List of lower taxa


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